Günter Bechly

Günter Bechly (1963-2025), German entomologist and paleontologist

• Biologic Institute, Redmond, Washington, USA
• Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany. (1999–2016)

(List may be incomplete)

• 90 taxon names authored by Günter Bechly

(List may be incomplete)

• 1996. Morphologische Untersuchungen am Flugelgeäder der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Berücksichtigung der phylogenetischen Systematik und des Grundplanes der Odonata. Petalura Special Volume 2: 402 pp, 3 tabls, 111 figs. Reference page. 
• , Nel, A. & Martínez Delclòs, X. 1996. Redescription of Nannogomphus bavaricus Handlirsch, 1906-1908 from the Upper Jurassic of Germany, with an analysis of its phylogenetic position (Odonata: Anisoptera: Gomphidae or Libelluloidea). Archaeopteryx 14: 51–66. PDF. Reference page. 

• 1997. New fossil odonates from the Upper Triassic of Italy, with a redescription of Italophlebia gervasuttii Whalley, and a reclassification of Triassic dragonflies (Insecta: Odonata). Rivista del Museo Civico di Scienze Naturali "Enrico Caffi" 19: 31–70. PDF. Reference page. 

• 1998. New fossil dragonflies from the Lower Cretaceous Santana Formation of north-east Brazil (Insecta: Odonata). Stuttgarter Beiträge zur Naturkunde, Serie B 264: 1–66. BHL. Reference page. 
• 1998. New fossil damselflies from Baltic amber, with description of a new species, a redescription of Litheuphaea carpenteri Fraser, and a discussion on the phylogeny of Epallagidae (Zygoptera: Caloptera). International Journal of Odonatology 1(1): 33–63. DOI: 10.1080/13887890.1998.9748092 . Reference page. 
• 1998. A revision of the fossil dragonfly genus Urogomphus, with description of a new species (Insecta: Odonata: Pananisoptera: Aeschnidiidae). Stuttgarter Beiträge zur Naturkunde, Serie B 270: 1–47. BHL. Reference page. 
• 1998. Juracordulia schiemenzi gen. et. sp. nov., Eine neue Libelle aus den Solnhofener Plattenkalken (Insecta: Odonata: Anisoptera). Archaeopteryx 16: 29–36. PDF. Reference page. 
• , Nel, A., Martínez-Delclòs, X. & Fleck, G. 1998. Four new dragonfly species from the Upper Jurassic of Germany and the Lower Cretaceous of Mongolia (Anisoptera: Hemeroscopidae, Sonidae, and Proterogomphidae fam. nov.). Odonatologica 27(2): 149–187. PDF. Reference page. 

• 2000. Two new fossil dragonfly species (Insecta: Odonata: Pananisoptera: Aeschnidiidae and Aktassiidae) from the Solnhofen Lithographic Limestones (Upper Jurassic of Germany). Stuttgarter Beiträge zur Naturkunde, Serie B 288: 1–9. BHL. Reference page. 
• 2000. Two new fossil dragonfly species (Insecta: Odonata: Anisoptera: Araripegomphidae and Lindeniidae) from the Crato Limestone (Lower Cretaceous, Brazil). Stuttgarter Beiträge zur Naturkunde, Serie B 296: 1–16. BHL. Reference page. 
• 2000. A new fossil damselfly species (Insecta: Odonata: Zygoptera: Coenagrionidae: Ischnurinae) from Dominican Amber. Stuttgarter Beiträge zur Naturkunde, Serie B 299: 1–9. BHL. Reference page. 
• & Schweigert, G. 2000. The first fossil hanging flies (Insecta: Mecoptera: Raptipedia: Cimbrophlebiidae and Bittacidae) from the limestones of Solnhofen and Nusplingen (Upper Jurassic of Germany). Stuttgarter Beiträge zur Naturkunde, Serie B 287: 1–18. PDF. Reference page. 

• 2001. A new species of Cymatophlebia (Insecta: Odonata: Anisoptera: Cymatophlebiidae) from the Solnhofen Lithographic Limestone (Upper Jurassic, Germany). Stuttgarter Beiträge zur Naturkunde, Serie B 301: 1–5. BHL. Reference page. 
• , Nel, A., Martínez-Delclòs, X., Jarzembowski, E.A., Coram, R., Martill, D., Fleck, G., Escuillié, F., Wisshak, M.M. & Maisch, M. 2001. A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Paläontologische Abhandlungen 4: 1–219. ISBN 3-931689-07-7. ResearchGate . Reference page. 
• , Brauckmann, C., Zessin, W. & Gröning, E. 2001. New results concerning the morphology of the most ancient dragonflies (Insecta: Odonatoptera) from the Namurian of Hagen-Vorhalle (Germany). Journal of Zoological Systematics and Evolutionary Research 39(4): 209–226. DOI: 10.1046/j.1439-0469.2001.00165.x . ResearchGate . Reference page. 
• Schawaller, W. & 2001. Die Ordnung der Vielfalt. In: Bechly, G. Ur-Geziefer. Die faszinierende Evolution der Insekten. Stuttgarter Beiträge zur Naturkunde (C) 49: 65–81. Full volume (PDF). Reference page.  

• & Sach, V.J. 2002. An interesting new fossil dragonfly (Anisoptera: Libellulidae: “Brachydiplacini”) from the Miocene of Germany, with a discussion on the phylogeny of Tetrathemistinae and a fossil list for the locality Heggbach. Stuttgarter Beiträge zur Naturkunde, Serie B 325: 1–11. PDF. Reference page. 
• & Ueda, K. 2002. The first fossil record and first New World record for the dragonfly clade Chlorogomphida (Insecta: Odonata: Anisoptera: Araripechlorogomphidae n. fam.) from the Crato Limestone (Lower Cretaceous, Brazil). Stuttgarter Beiträge zur Naturkunde, Serie B 328: 1–11. PDF. Reference page. 
• Huguet, A., Nel, A., Martínez-Delclòs, X., & Martins-Neto, R. 2002. Preliminary phylogenetic analysis of the Protanisoptera (Insecta: Odonatoptera). Geobios 35(5): 537–560. DOI: 10.1016/S0016-6995(02)00071-2 . ResearchGate . Reference page. 
• Nel, A., Béthoux, O., , Martínez-Delclòs, X. & Papier, F. 2002 ("2001"). The Permo-Triassic Odonatoptera of the ‘protodonate’ grade (Insecta: Odonatoptera). Annales de la Société Entomologique de France 37(4): 501–525. Gallica, Academia.edu, ResearchGate . Reference page. 

• 2003. Description of a new species of Nannogomphus (Insecta: Odonata: Nannogomphidae) from the Upper Jurassic Solnhofen Limestone in Germany. Stuttgarter Beiträge zur Naturkunde, Serie B 339: 1–6. PDF. Reference page. 
• 2003. Odonatology Website [offline version 2002]. In: Schorr, M. & Lindeboom, M. (eds.) Dragonfly Research 1.2003. Zerf-Tübingen (CD-ROM). Reference page. 
• , Dietl, G. & Schweigert, G. 2003. A new species of Stenophlebia (Insecta: Odonata: Stenophlebiidae) from the Nusplingen Lithographic Limestone (Upper Jurassic, SW Germany). Stuttgarter Beiträge zur Naturkunde, Serie B 338: 1–10. PDF. Reference page. 
• Fleck, G., , Martínez-Delclòs, X., Jarzembowski, E., Coram, R. & Nel, A. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Société entomologique de France (n.s.) 39(1): 55–93. DOI: 10.1080/00379271.2003.10697363 . Reference page. 

• 2005. A new fossil dragonfly (Anisoptera: Corduliidae) from the Paleocene Fur Formation (Mo clay) of Denmark. Stuttgarter Beiträge zur Naturkunde, Serie B 358: 1–7. PDF. Reference page. 
• 2005. A re-description of "Stenophlebia" casta (Insecta: Odonata: Parastenophlebiidae n. fam) from the Upper Jurassic Solnhofen Limestone in Germany. Stuttgarter Beiträge zur Naturkunde, Serie B 359: 1–12. PDF. Reference page. 
• Nel, A., , Garrouste, R., Pohl, B. & Escuillié, F. 2005. A new extraordinary neuropterid family from the Lower Cretaceous Crato Formation of Brazil: a new insect order? (Insecta, Neuropterida). Cretaceous Research 26(6): 845–852. DOI: 10.1016/j.cretres.2005.05.006 Reference page. 

• 2007. Isoptera: termites. In D. M. Martill, G. Bechly, R. F. Loveridge (eds.), The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, 624 pp. ISBN 9780521858670 Reference page. 
• 2007. Odonata: damselflies and dragonflies. In D. M. Martill, G. Bechly, R. F. Loveridge (eds.), The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, 624 pp. ISBN 9780521858670 Reference page. 
• 2007. Blattaria: cockroaches and roachoides. In D. M. Martill, G. Bechly, R. F. Loveridge (eds.), The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, 624 pp. ISBN 9780521858670 Reference page. 

• Delclòs, X., Azar, A., , Dunlop, J.A., Engel, M.S. & Heads, S.W. 2008. The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): New palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 247(3): 353–381. DOI: 10.1127/0077-7749/2008/0247-0353 Full article (PDF) on ResearchGate Reference page. 
• Fleck, G., Nel, A., , Delclòs, X., Jarzembowski, E.A. & Coram, R. 2008. New Lower Cretaceous 'libelluloid' dragonflies (Insecta: Odonata: Cavilabiata) with notes about estimated divergence dates for this group. Palaeodiversity 1: 19–36. PDF. Reference page. 

• 2010. Additions to the fossil dragonfly fauna of the Lower Cretaceous Crato Formation of Brazil (Insecta: Odonata). Palaeodiversity 3 (Supplement "Contributions to the WILLI-HENNIG-Symposium on Phylogenetics and Evolution, University of Hohenheim, 29 September - 2 October 2009"): 11–77. PDF. Reference page. 
• Poinar, G.O., jr., & Buckley, R. 2010. First record of Odonata and a new subfamily of damselflies from Early Cretaceous Burmese amber. Palaeodiversity 3: 15–22. PDF. Reference page. 

• & Stockar, R. 2011. The first Mesozoic record of the extinct apterygote insect genus Dasyleptus (Insecta: Archaeognatha: Monura: Dasyleptidae) from the Triassic of Monte San Giorgio (Switzerland). Palaeodiversity 4: 23–37. PDF. Reference page. 
• & Wolf-Schwenninger, K. 2011. A new fossil genus and species of snakefly (Raphidioptera: Mesoraphidiidae) from Lower Cretaceous Lebanese amber, with a discussion of snakefly phylogeny and fossil history. Insect Systematics & Evolution 42(2): 221–236. DOI: 10.1163/187631211X568164 Reference page. 
• Staniczek, A.H., & Godunko, R.J. 2011. Coxoplectoptera, a new fossil order of Palaeoptera (Arthropoda: Insecta), with comments on the phylogeny of the stem group of mayflies (Ephemeroptera). Insect Systematics & Evolution 42(2): 101–138. DOI: 10.1163/187631211X578406 . PDF. Reference page. 

• 2012. An interesting new fossil relict damselfly (Odonata: Zygoptera: Coenagrionoidea) from Eocene Baltic amber. Palaeodiversity 5: 51–55. PDF. Reference page. 
• Nel, A., , Prokop, J., Béthoux, O. & Fleck, G. 2012. Systematics and evolution of Paleozoic and Mesozoic damselfly-like Odonatoptera of the “protozygopteran” grade. Journal of Paleontology 86(1): 81–104. DOI: 10.1666/11-020.1 . Reference page. 

• & Kin, A.(†) 2013. First record of the fossil dragonfly family Eumorbaeschnidae (Insecta: Odonata: Anisoptera) from the Upper Jurassic of Poland. Acta Palaeontologica Polonica 58(1): 121–124. DOI: 10.4202/app.2011.0194 . Reference page. 
• & Poinar, G., Jr. 2013. Burmaphlebia reifi gen. et sp. nov., the first anisozygopteran damsel-dragonfly (Odonata: Epiophlebioptera: Burmaphlebiidae fam. nov.) from Early Cretaceous Burmese amber. Historical Biology 25(2): 233–237. DOI: 10.1080/08912963.2012.753884 . PDF. Reference page. 
• Dijkstra, K.-D.B., , Bybee, S.M., Dow, R.A., Dumont, H.J., Fleck, G., Garrison, R.W., Hämäläinen, M., Kalkman, V.J., Karube, H., May, M.L., Orr, A.G., Paulson, D.R., Rehn, A.C., Theischinger, G., Trueman, J.W.H., van Tol, J., von Ellenrieder, N. & Ware, J. 2013. The classification and diversity of dragonflies and damselflies (Odonata). In: Zhang, Z.-Q. (ed.) 2013. Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness (Addenda 2013). Zootaxa 3703(1): 36–45. DOI: 10.11646/zootaxa.3703.1.9 . Reference page. 
• Rasser, M.W., , Böttcher, R., Ebner, M., Heizmann. E.P.J., Höltke, O., Joachim, C., Kern, A.K., Kovar-Eder, J., Nebelsick, J.H., Roth-Nebelsick, A., Schoch, R.R., Schweigert, G. & Ziegler, R. 2013. The Randeck Maar: Palaeoenvironment and habitat differentiation of a Miocene lacustrine system. Palaeogeography, Palaeoclimatology, Palaeoecology 392: 426–453. DOI: 10.1016/j.palaeo.2013.09.025 Reference page. 

• Greenwalt, D.E. & 2014. A re-description of the fossil damselfly Eolestes syntheticus Cockerell, 1940 (Odonata: Zygoptera: Eolestidae n. fam.) with description of new taxa from the Eocene of North America. Zootaxa 3887(2): 138–156. DOI: 10.11646/zootaxa.3887.2.2 . PDF. Reference page. 
• Sroka, P., Staniczek, A.H. & 2014. Revision of the giant pterygote insect Bojophlebia prokopi Kukalová-Peck, 1985 (Hydropalaeoptera: Bojophlebiidae) from the Carboniferous of the Czech Republic, with the first cladistic analysis of fossil palaeopterous insects. Journal of Systematic Palaeontology 13(11): 963–982. DOI: 10.1080/14772019.2014.987958 . Reference page. 
• Staniczek, A.H., Sroka, P. & 2014. Neither silverfish nor fowl: the enigmatic Carboniferous Carbotriplura kukalovae Kluge, 1996 (Insecta: Carbotriplurida) is the putative fossil sister group of winged insects (Insecta: Pterygota). Systematic Entomology 39(4): 619–632. DOI: 10.1111/syen.12076 . Reference page. 

• Caterino, M.S., Wolf-Schwenninger, K. & 2015. Cretonthophilus tuberculatus, a remarkable new genus and species of hister beetle (Coleoptera: Histeridae) from Cretaceous Burmese amber. Zootaxa 4052(2): 241–245. DOI: 10.11646/zootaxa.4052.2.10. Preview (PDF) Full article (PDF) Reference page. 
• Dunlop, J.A., Bird, T.L., Brookhart, O. & 2015. A camel spider from Cretaceous Burmese amber. Cretaceous Research 56: 265–273. DOI: 10.1016/j.cretres.2015.05.003 Reference page. 

• & Makarkin, V.N. 2015 (online) 2016 (print). A new giantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the New World. Cretaceous Research 58: 135–140. DOI: 10.1016/j.cretres.2015.10.014 . PDF. Reference page. 
• Huang, D.-Y., , Nel, P., Engel, M.S., Prokop, J., Azar, D., Cai, C.-Y., van de Kamp, T., Staniczek, A.H., Garrouste, R., Krogmann, L., dos Santos Rolo, T., Baumbach, T., Ohlhoff, R., Shmakov, A.S., Bourgoin, T. & Nel, A. 2016. New fossil insect order Permopsocida elucidates major radiation and evolution of suction feeding in hemimetabolous insects (Hexapoda: Acercaria). Scientific Reports 6: 23004. DOI: 10.1038/srep23004 . Reference page. 
• Schädel, M. & 2016. First Record of Anisoptera (Insecta: Odonata) from mid-Cretaceous Burmese Amber. Zootaxa 4103(6): 537–549. DOI: 10.11646/zootaxa.4103.6.4. Reference page. 

• Möstel, C., Schorr, M. & 2017. A new stem-coenagrionoid genus of damselflies (Odonata: Zygoptera) from mid-Cretaceous Burmese amber. Zootaxa 4243(1): 177–186. DOI: 10.11646/zootaxa.4243.1.9 Reference page. 
• Pinkert, S., & Nel, A. 2017. First record of hawker dragonflies from Eocene Baltic amber (Odonata: Anisoptera: Gomphaeschnidae). Zootaxa 4272(2): 263–275. DOI: 10.11646/zootaxa.4272.2.7 . Reference page. 

• 2018. First record and a new species of the fossil dragonfly genus Proinogomphus (Odonata: Liassogomphidae) from the Early Jurassic of Bascharage in the Grand Duchy of Luxembourg. Zootaxa 4450(1): 108–114. DOI: 10.11646/zootaxa.4450.1.7 . PDF. Reference page. 
• 2018. Chrismooreia michaelbehei gen. et sp. nov. (Insecta: Odonata: Asiopteridae), a new fossil damsel-dragonfly from the Early Jurassic of England. BIO-Complexity 2018(1): 1–10. DOI: 10.5048/BIO-C.2018.1 . Reference page. 
• Vršanský, P., , Zhang, Q., Jarzembowski, E.A., Mlynský, T., Šmídová, L., Barna, P., Kúdela, M., Aristov, D., Bigalk, S., Krogmann, L., Li, L., Zhang, Q., Zhang, H., Ellenberger, S., Müller, P., Gröhn, C., Xia, F., Ueda, K., Vďačný, P., Valaška, D., Vršanská, L. & Wang, B. 2018. Batesian insect-insect mimicry-related explosive radiation of ancient alienopterid cockroaches. Biologia 73(10): 987–1006. DOI: 10.2478/s11756-018-0117-3 . PDF. Reference page. 

• 2019. New fossil Odonata from the Upper Jurassic of Bavaria with a new fossil calibration point for Zygoptera. Palaeoentomology 2(6): 618–632. DOI: 10.11646/palaeoentomology.2.6.13 . PDF. Reference page. 
• & Rasmussen, J.A. 2019. A new genus of hawker dragonfly (Odonata: Anisoptera: Aeshnidae) from the Early Eocene Fur Formation of Denmark. Zootaxa 4550(1): 123–128. DOI: 10.11646/zootaxa.4550.1.6 . Plazi. Reference page. 
• Vršanský, P., Sendi, H., Aristov, D., , Müller, P., Ellenberger, S., Azar, D., Ueda, K., Barna, P. & Garcia, T. 2019. Ancient roaches further exemplify ‘no land return’ in aquatic insects. Gondwana Research 68: 22–33. DOI: 10.1016/j.gr.2018.10.020 . PDF. Reference page. 

• , Garrouste, R., Aase, A., Karr, J.A., Grande, L. & Nel, A. 2020 (Online) 2021 (Print). The damselfly palaeofauna from the Eocene of Wyoming and Colorado, USA (Insecta, Odonata, Zygoptera). Papers in Palaeontology 7(3): 1373–1402. DOI: 10.1002/spp2.1346 . Reference page. 

• & Velten, J. 2023. A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263(4): 547–556. DOI: 10.11646/zootaxa.5263.4.6 . Plazi. Reference page. 

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