Grylloblattodea

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Kingdom: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Mandibulata
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Supercohort: Polyneoptera
Ordo:
Familiae: Grylloblattidae

Ordo Grylloblattida secundum Storozhenko (1997, 1998)
Subordines: Grylloblattina – †Lemmatophorina – †Protoperlina

Unplaced to suborder: †Aristoviidae – †Sinonamuropteridae

Grylloblattida incertae sedis: †Cabarzopterum – †Permedax

Taxon Blattogryllopterida secundum Cui (2012) + Cui et al. (2024)
Familiae: Grylloblattidae – †Blattogryllidae
Genera: †Aristovia – †Aristoviblattogryllus

Ordo Eoblattida secundum Aristov (2015)
Familiae: Grylloblattidae – †Atactophlebiidae – †Bardapteridae – †Blattogryllidae – †Cacurgidae – †Daldubidae – †Doubraviidae – †Eoblattidae – †Euryptilonidae – †Ideliidae – †Idelinellidae – †Megakhosaridae – †Mesorthopteridae – †Permopectinidae – †Permotermopsidae – †Protophasmatidae – †Soyanopteridae – †Tococladidae

Ordo Reculida secundum Aristov (2015)
†Archiprobnidae – †Bajanzhargalanidae – †Camptoneuritidae – †Chaulioditidae – †Chelopteridae – †Epideigmatidae – †Euremiscidae – †Geinitziidae – †Gorochoviidae – †Havlatiidae – †Ivapteridae – †Kargalopteridae – †Lemmatophoridae – †Liomopteridae – †Madygenophlebidae – †Neleidae – †Probnidae – †Protoblattinidae – †Reculidae – †Sinonamuropteridae – †Skaliciidae – †Sylvaphlebiidae

Grylloblattodea Brues & Melander, 1932

• Grylloblattoidea Brues & Melander, 1915: 13
• Grylloblattodea Brues & Melander, 1932: 47
• Grylloblattida

• Brues, C. & Melander, A. 1915. Key to the families of North American insects; an introduction to the classification of insects. Boston, Massachusetts and Pullman, Washington. 140 pp., 18 pls. DOI: 10.5962/bhl.title.19662 . BHL. Reference page. 
• Brues, C. & Melander, A. 1932. Classification of Insects. A key to the known families of insects and other terrestrial arthropods. Bulletin of the Museum of Comparative Zoology at Harvard College 73: 672 pp., 1121 figs. BHL. Reference page. 

• Cui, Y., Bardin, J., Wipfler, B., Demers-Potvin, A., Bai, M., Tong, Y.-J., Chen, G.N., Chen, H., Zhao, Z.-Y., Ren, D. & Béthoux, O. 2024. A winged relative of ice-crawlers in amber bridges the cryptic extant Xenonomia and a rich fossil record. Insect Science 31(5): 1645–1656. DOI: 10.1111/1744-7917.13338 . Reference page.  [proposes new unranked taxa Icaroptera and Tenuivenaptera]
• Jarvis, K.J. 2005. Phylogeny and biogeography of ice crawlers (Insecta: Grylloblattodea): evidence from six molecular loci. UNPUBLISHED M.SC. THESIS, BRIGHAM YOUNG UNIVERSITY, UTAH, U.S.A. PDF.
• Jarvis, K.J. & Whiting, M.F. 2006. Phylogeny and biogeography of ice crawlers (Insecta: Grylloblattodea) based on six molecular loci: designating conservation status for Grylloblattodea species. Molecular Phylogenetics and Evolution 41(1): 222–237. DOI: 10.1016/j.ympev.2006.04.013 . Reference page. 
• Schoville, S.D. 2019. Grylloblattodea of Canada. Pp 271–276 In Langor, D.W. & Sheffield, C.S. (eds.). The Biota of Canada – A Biodiversity Assessment. Part 1: The Terrestrial Arthropods. ZooKeys 819: 520 pp. Reference page. . DOI: 10.3897/zookeys.819.24289 Reference page. 
• Walker, E.M. 1914. A new species of Orthoptera, forming a new genus and family. The Canadian Entomologist 46: 93–99. BHL.
• Wipfler, B., Bai, M., Schoville, S., Dallai, R., Uchifune, T., Machida, R., Cui, Y. & Beutel, R.G. 2014. Ice Crawlers (Grylloblattodea) – the history of the investigation of a highly unusual group of insects. Journal of Insect Biodiversity 2(2): 1–25. DOI: 10.12976/jib/2014.2.2 . Reference page. 

Grylloblattida

• Aristov, D.S. 2009. [A new family of the order Grylloblattida (Insecta) from the Middle Permian of Russia]. Paleontologicheskii Zhurnal 2009(2): 59–63. (in Russian). [English translation in Paleontological Journal 43(2): 178–182. DOI: 10.1134/S0031030109020087 . ResearchGate .] Reference page. 
• Aristov, D.S. & Rasnitsyn, A.P. 2010. The family Tillyardembiidae Zalessky, 1938 and the system of the plecopteroid insects. Russian Entomological Journal 18(4) (for 2009): 257–264. PDF. Reference page. 
• Béthoux, O., Nel, A., Lapeyrie, J. & Gand, G. 2005. New data on Paleozoic grylloblattid insects (Neoptera). Journal of Paleontology 79(1): 125–145. DOI: 10.1666/0022-3360(2005)079<0125:NDOPGI>2.0.CO;2 . [Errata: 79(3): 629–630. DOI: 10.1666/0022-3360(2005)079<0629:E>2.0.CO;2 . JSTOR .] Reference page. 
• Cui, Y. 2012. New data on the Blattogryllidae-Plesioblattogryllidae-Grylloblattidae complex (Insecta: Grylloblattida: Blattogryllopterida tax.n.). Arthropod Systematics & Phylogeny 70(3): 167–180. PDF. Reference page.  [proposes new unranked taxon Blattogryllopterida]
• Cui, Y., Béthoux, O. & Ren, D. 2011. Intraindividual variability in Sinonamuropteridae forewing venation (Grylloblattida; Late Carboniferous): taxonomic and nomenclatural implications. Systematic Entomology 36(1): 44–56. DOI: 10.1111/j.1365-3113.2010.00545.x . Reference page. 
• Cui, Y., Brauner, S., Schneider, J.W. & Béthoux, O. 2021. Grylloblattidan insects from Sperbersbach and Cabarz (Germany), two new early Permian and insect-rich localities. Journal of Paleontology 96(2): 355–374. DOI: 10.1017/jpa.2021.101 . Reference page. 
• Cui, Y., Béthoux, O., Yang, N. & Ren, D. 2023. New data and systematic considerations on the Blattogryllopterida (Insecta: Grylloblattida) from the Jurassic Daohugou locality in China. Palaeoentomology 6(2): 124–132. DOI: 10.11646/palaeoentomology.6.2.4 . Plazi. Reference page. 
• Huang, D.Y., Nel, A. & Petrulevičius, J.F. 2008. New evolutionary evidence of Grylloblattida from the Middle Jurassic of Inner Mongolia, north-east China (Insecta, Polyneoptera). Zoological Journal of the Linnean Society 152(1): 17–24. DOI: 10.1111/j.1096-3642.2007.00351.x . Reference page. 
• Rasnitsyn, A.P. & Quicke, D.L.J. (eds.) 2002. History of Insects. Kluwer Academic Publishers, New York, Boston, Dordrecht, London, Moscow, 517 pp. ISBN 1-4020-0026-X (print), ISBN 0-306-47577-4 (e-book). Online version. DOI: 10.1007/0-306-47577-4 . Reference page. 
• Storozhenko, S.Yu. 1997. Classification of the order Grylloblattida (Insecta), with description of new taxa. Far Eastern Entomologist 42: 1–20. PDF.
• Storozhenko, S.Yu. 1998. Систематика, филогения и эволюция гриллоблаттидовых насекомых [Systematics, phylogeny and evolution of the grylloblattids (Insecta: Grylloblattida)]. Vladivostok: Dal'nauka. 207 pp. (in Russian). PDF.
• Storozhenko, S.Yu. 2002. 2.2.2.2.1. Order Grylloblattida Walker, 1914 (=Notoptera Crampton, 1915, =Grylloblattodea Brues & Melander, 1932, +Protorthoptera Handlirsch, 1906, =Paraplecoptera Martynov, 1925, +Protoperlaria Tillyard, 1928). Pp. 278–281. In: Rasnitsyn, A.P. & Quicke, D.L.J. (eds.) History of Insects. Kluwer Academic Publishers, New York, Boston, Dordrecht, London, Moscow. ISBN 1-4020-0026-X (print), ISBN 0-306-47577-4 (e-book). DOI: 10.1007/0-306-47577-4 . Reference page. 
• Storozhenko, S.Yu. & Gröhn, C. 2023. A new family of grylloblattids (Insecta: Grylloblattida) from mid-Cretaceous Burmese amber. Palaeoentomology 6(2): 165–173. DOI: 10.11646/palaeoentomology.6.2.8 . ResearchGate . FSC EATB FEB RAS (PDF) Reference page. 

Eoblattida/Reculida

• Aristov, D.S. 2015. Classification of the order Eoblattida (Insecta: Blattidea) with description of new taxa. Far Eastern Entomologist 301: 1–56. PDF.
• Rasnitsyn, A.P. & Aristov, D.S. 2021. Обзор семейства Tococladidae Carpenter (Insecta: Eoblattida) из перми Северной Америки и Европы. [Review of the family Tococladidae Carpenter (Insecta: Eoblattida) from the Permian of North America and Europe.] Paleontologicheskii Zhurnal 2021(2): 37–41. (In Russian). [English translation: Paleontological Journal 55(2): 151–156. DOI: 10.1134/S003103012102012X . ResearchGate .] Reference page. 

• Grylloblattodea Species File

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