Gustavo Hormiga

Gustavo A. Hormiga, US arachnologist

• Department of Biological Sciences, The George Washington University, Washington, DC 20052.
• e-mail: hormiga@gwu.edu

(List may be incomplete)

• 106 taxon names authored by Gustavo Hormiga

(List may be incomplete)

• 1994. A revision and cladistic analysis of the spider family Pimoidae (Araneoidea: Araneae). Smithsonian Contributions to Zoology 549: 1-104. Reference page. 

• Griswold, C.E., Long, C.L. & 1999. A new spider of the genus Pimoa from Gaoligong Mountains, Yunnan, China (Araneae, Araneoidea, Pimoidae). Acta Botanica Yunnanica Supplement 11: 91-97. Reference page. 

• ; Buckle, D.J. & Scharff, N 2005: Nanoa, an enigmatic new genus of pimoid spiders from western North America (Pimoidae, Araneae). Zoological journal of the Linnean Society, 145: 249–262. DOI: 10.1111/j.1096-3642.2005.00192.x Reference page. 
• & Scharff, N. 2005. Monophyly and phylogenetic placement of the spider genus Labulla Simon, 1884 (Araneae, Linyphiidae) and description of the new genus Pecado. Zoological journal of the Linnean Society 143(3): 359–404. DOI: 10.1111/j.1096-3642.2005.00147.x Reference page. 

• , Alvarez-Padilla, F. & Benjamin, S.P. 2007. First records of extant Hispaniolan spiders of the families Mysmenidae, Symphytognathidae, and Ochyroceratidae (Araneae), including a new species of Ochyrocera. American Museum Novitates 3577: 1-21. Reference page. 
• Lopardo, & Melic, A. 2007. Spinneret spigot morphology in synaphrid spiders (Araneae, Synaphridae), with comments on the systematics of the family and description of a new species of Synaphris Simon 1894 from Spain. American Museum Novitates 3556: 1-16. Reference page. 

• 2008. On the spider genus Weintrauboa (Araneae, Pimoidae), with a description of a new species from China and comments on its phylogenetic relationships. Zootaxa 1814: 1-20. DOI: 10.11646/zootaxa.1814.1.1 Reference page. 
• Kuntner, M., Coddington, J.A. & 2008. Phylogeny of extant nephilid orb-weaving spiders (Araneae, Nephilidae): testing morphological and ethological homologies. Cladistics 24: 190. DOI: 10.1111/j.1096-0031.2007.00176.x Reference page. 

• Tu, L.H. & 2011. Phylogenetic analysis and revision of the linyphiid spider genus Solenysa (Araneae: Linyphiidae: Erigoninae). Zoological Journal of the Linnean Society 161: 484-530. Reference page. 

• Mammola, S., , Arnedo, M.A. & Isaia, M. 2016. Unexpected diversity in the relictual European spiders of the genus Pimoa (Araneae : Pimoidae). Invertebrate Systematics 30: 566–587. Reference page. 
• Scharff, N. & 2012: First evidence of aggressive chemical mimicry in the Malagasy orb weaving spider Exechocentrus lancearius Simon, 1889 (Arachnida: Araneae: Araneidae) and description of a second species in the genus. Arthropod systematics & phylogeny, 70(2): 107–118. PDF.Reference page. 

• Andriamalala, D. & 2013. Systematics of the goblin spider genus Opopaea (Araneae, Oonopidae) in Magagascar. Bulletin of the American Museum of Natural History 380: 1-156. DOI: 10.1206/828.1 . hdl: 2246/6443 . Reference page. 
• Scharff, N. & 2013: On the Australian linyphiid spider Alaxchelicera ordinaria Butler, 1932 (Araneae). Zootaxa 3750(2): 193–196. DOI: 10.11646/zootaxa.3750.2.8 Reference page. 

• Lopardo & 2015. Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea). Zoological Journal of the Linnean Society 173(3): 527-786. DOI: 10.1111/zoj.12199. Reference page. 
• Silva-Moreira, T. da & 2015. Redescription and phylogenetic placement of the Hispaniolan spider genus Lomaita Bryant, 1948 (Araneae, Linyphiidae). Zootaxa 3920(2): 249–264. DOI: 10.11646/zootaxa.3920.2.2. Reference page. 

• Benavides, L.R. & 2016. Taxonomic revision of the Neotropical pirate spiders of the genus Gelanor Thorell, 1869 (Araneae, Mimetidae) with the description of five new species. Zootaxa 4064(1): 1–72. DOI: 10.11646/zootaxa.4064.1.1. Reference page. 
• Kallal, R.J. & 2016. Redescription of the Jamaican orb-weaving spider Chrysometa pecki Levi, 1986 (Araneae, Tetragnathidae), including the first description of the male. Zootaxa 4066(1): 95–100. DOI: 10.11646/zootaxa.4066.1.11. Reference page. 

• 2017. The discovery of the orb-weaving spider genus Pinkfloydia (Araneae, Tetragnathidae) in eastern Australia with description of a new species from New South Wales and comments on the phylogeny of Nanometinae. Zootaxa 4311(4): 480–490. DOI: 10.11646/zootaxa.4311.4.2. Reference page. 
• & Dimitrov, D. 2017. The discovery of the spider genus Putaoa (Araneae, Pimoidae) in Taiwan with the description of a new species, including its web architecture. Zootaxa 4341(1): 97–104. DOI: 10.11646/zootaxa.4341.1.8. Reference page. 

• Kallal, R.J. & 2018. An expanded molecular phylogeny of metaine spiders (Araneae, Tetragnathidae) with description of new taxa from Taiwan and the Philippines. Invertebrate Systematics 32(2): 400-422. DOI: 10.1071/IS17058. Reference page. 
• Kallal, R.J. & 2018. Systematics, phylogeny and biogeography of the Australasian leaf-curling orb-weaving spiders (Araneae: Araneidae: Zygiellinae), with a comparative analysis of retreat evolution. Zoological Journal of the Linnean Society 184: 1055–1141. DOI: 10.1093/zoolinnean/zly014 Reference page. 

• Benavides, L.R., & Giribet, G. 2019. Phylogeny, evolution and systematic revision of the mite harvestman family Neogoveidae (Opiliones Cyphophthalmi). Invertebrate Systematics 33(1): 101–180. DOI: 10.1071/IS18018 Reference page. 
• Kallal, R.J. & 2019. Evolution of the male palp morphology of the orb-weaver hunting spider Chorizopes (Araneae : Araneidae) revisited on a new phylogeny of Araneidae, and description of a third species from Madagascar. Invertebrate Systematics 33(2): 473-487. DOI: 10.1071/IS18061. Reference page. 

• Alvarez-Padilla, F., Kallal, R.J. & 2020. Taxonomy and phylogenetics of Nanometinae and other Australasian orb-weaving spiders (Araneae: Tetragnathidae). Bulletin of the American Museum of Natural History 438: 1-107. DOI: 10.1206/0003-0090.438.1.1. . Reference page. 
• Benavides, L.R. & 2020. A morphological and combined phylogenetic analysis of pirate spiders (Araneae, Mimetidae): evolutionary relationships, taxonomy, and character evolution. Invertebrate Systematics 34(2): 144-191. DOI: 10.1071/IS19032 . Reference page. 
• & Scharff, N. 2020. The malkarid spiders of New Zealand (Araneae : Malkaridae). Invertebrate Systematics 34(4): 345–405. DOI: 10.1071/IS19073. Reference page. 

• Arnedo, M.A. & 2021. Repeated colonization, adaptive radiation and convergent evolution in the sheet‐weaving spiders (Linyphiidae) of the south Pacific Archipelago of Juan Fernandez. Cladistics 37(3): 317-342. DOI: 10.1111/cla.12437. . Reference page. 
• Ballesteros, J.A. & 2021. Molecular phylogeny of the orb-weaving spider genus Leucauge and the intergeneric relationships of Leucauginae (Araneae, Tetragnathidae). Invertebrate Systematics 35(8): 922-939. DOI: 10.1071/IS21029. . Reference page. 
• Hazzi, N.A. & 2021. Morphological and molecular evidence support the taxonomic separation of the medically important Neotropical spiders Phoneutria depilata (Strand, 1909) and P. boliviensis (F.O. Pickard-Cambridge, 1897) (Araneae, Ctenidae). Zookeys 1022: 13–50. DOI: 10.3897/zookeys.1022.60571 Reference page. 
• Silva-Moreira, T. da & 2021. Systematics of the Neotropical spider genera Jalapyphantes and Selenyphantes and the circumscription of the Pocobletus clade (Araneae: Linyphiidae) Zoological Journal of the Linnean Society 192(3): 896–957. DOI: 10.1093/zoolinnean/zlaa124 Reference page. 

• Kallal, R.J. & 2020. Phylogenetic placement of the stone-nest orb-weaving spider Nemoscolus Simon, 1895 (Araneae : Araneidae) and the description of the first species from Australia. Invertebrate Systematics 34(8): 893-905. DOI: 10.1071/IS20035. Reference page. 
• Silva-Moreira, T. da & 2022. Revision and phylogenetics of the Neotropical sheet weaving spider genus Diplothyron Millidge, 1991 (Araneae, Linyphiidae) and systematics of the MPME clade. Invertebrate Systematics 36(9): 781-848. DOI: 10.1071/IS21047 Reference page. 

• Hazzi, N.A. & 2023. Molecular phylogeny of the tropical wandering spiders (Araneae, Ctenidae) and the evolution of eye conformation in the RTA clade. Cladistics 39(1): 18-42. DOI: 10.1111/cla.12518. . Reference page. 
• Liu, J.X., Xu, X., , Yin, H.Q. & Li, H. 2023. Two new species of the spider genus Putaoa (Araneae, Linyphiidae) from southern China, Zootaxa 5277(3): 553–564. DOI: 10.11646/zootaxa.5277.3.7. . Reference page. 

• Hazzi, N. & 2024. Systematics, distribution patterns and historical biogeography of the Central America wandering spider genus Kiekie Polotow & Brescovit, 2018 (Araneae: Ctenidae). Peerj DOI: 10.7717/peerj.17242. Reference page. 

• Hazzi, N.A., Wood, H.M. & 2025. Reassessing the evolutionary relationships of tropical wandering spiders using phylogenomics: A UCE-based phylogeny of Ctenidae (Araneae) with the discovery of a new lycosoid family. Molecular Phylogenetics and Evolution 203 08245. Reference page. 

• CV