Mark D. Scherz

Mark D. Scherz, German herpetologist.

• Zoologische Staatssammlung München, Münchhausenstraße 21, München, Germany.

(List may be incomplete)

• 139 taxon names authored by Mark D. Scherz

(List may be incomplete)

• , Ruthensteiner, B., Vences, M. & Glaw, F. 2014. A new microhylid frog, genus Rhombophryne, from northeastern Madagascar, and a re-description of R. serratopalpebrosa using micro-computed tomography. Zootaxa 3860(6): 547–560. DOI: 10.11646/zootaxa.3860.6.3 Reference page. 

• Prötzel, D., Ruthensteiner, B., & Glaw, F. 2015. Systematic revision of the Malagasy chameleons Calumma boettgeri and C. linotum (Squamata: Chamaeleonidae). Zootaxa 4048(2): 211–231. DOI: 10.11646/zootaxa.4048.2.4 Reference page. 
• , Ruthensteiner, B., Vieites, D.R., Vences, M. & Glaw, F. 2015. Two new microhylid frogs of the genus Rhombophryne with superciliary spines from the Tsaratanana Massif in northern Madagascar. Herpetologica 71(2): 310–321. DOI: 10.1655/HERPETOLOGICA-D-14-00048 Reference page. 
• , Rakotoarison, A., Hawlitschek, O., Vences, M. & Glaw, F. 2015. Leaping towards a saltatorial lifestyle? An unusually long-legged new species of Rhombophryne (Anura, Microhylidae) from the Sorata massif in northern Madagascar. Zoosystematics and Evolution 91(2): 105–114. DOI: 10.3897/zse.91.4979 Reference page. 

• Hawlitschek, O., , Straube, N. & Glaw, F. 2016. Resurrection of the Comoran fish scale gecko Geckolepis humbloti Vaillant, 1887 reveals a disjunct distribution caused by natural overseas dispersal. Organisms Diversity & Evolution 16(1): 289–298. DOI: 10.1007/s13127-015-0255-1 Reference page. 
• Lattenkamp, E.Z., Mandák, M. & 2016. The advertisement call of Stumpffia be Köhler, Vences, D’Cruze & Glaw, 2010 (Anura: Microhylidae: Cophylinae). Zootaxa 4205(5): 483–485. DOI: 10.11646/zootaxa.4205.5.7 Reference page. 
• , Vences, M., Rakotoarison, A., Andreone, F., Köhler, J., Glaw, F. & Crottini, A. 2016. Reconciling molecular phylogeny, morphological divergence and classification of Madagascan narrow-mouthed frogs (Amphibia: Microhylidae). Molecular Phylogenetics and Evolution 100: 372–381. DOI: 10.1016/j.ympev.2016.04.019 Reference page. 
• , Glaw, F., Vences, M., Andreone, F. & Crottini, A. 2016. Two new species of terrestrial microhylid frogs (Microhylidae: Cophylinae: Rhombophryne) from northeastern Madagascar. Salamandra 52(2): 91–106. PDF Reference page. 

• , Hawlitschek, O., Andreone, F., Rakotoarison, A., Vences, M. & Glaw, F. 2017. A review of the taxonomy and osteology of the Rhombophryne serratopalpebrosa species group (Anura: Microhylidae) from Madagascar, with comments on the value of volume rendering of micro-CT data to taxonomists. Zootaxa 4273(3): 301–340. DOI: 10.11646/zootaxa.4273.3.1 Reference page. 
• , Daza, J.D., Köhler, J., Vences, M. & Glaw, F. 2017. Off the scale: a new species of fish-scale gecko (Squamata: Gekkonidae: Geckolepis) with exceptionally large scales. PeerJ 5: e2955. DOI: 10.7717/peerj.2955 Reference page. 
• Lambert, S.M., Hutter, C.R. & 2017. Diamond in the rough: a new species of fossorial diamond frog (Rhombophryne) from Ranomafana National Park, southeastern Madagascar. Zoosystematics and Evolution 93(1): 143–155. DOI: 10.3897/zse.93.10188 Reference page. 
• Rakotoarison, A., , Glaw, F., Köhler, J., Andreone, F., Franzen, M., Glos, J., Hawlitschek, O., Jono, T., Mori, A., Ndriantsoa, S.H., Raminosoa, N.R., Riemann, J.C., Rödel, M.-O., Rosa, G.M., Vieites, D.R., Crottini, A. & Vences, M. 2017. Describing the smaller majority: integrative taxonomy reveals twenty-six new species of tiny microhylid frogs (genus Stumpffia) from Madagascar. Vertebrate Zoology 67(3): 271–398. PDF Reference page. 
• Ratsoavina, F.M., Gehring, P.-S., , Vieites, D.R., Glaw, F. & Vences, M. 2017. Two new species of leaf-tailed geckos (Uroplatus) from the Tsaratanana mountain massif in northern Madagascar. Zootaxa 4347(3): 446–464. DOI: 10.11646/zootaxa.4347.3.2 Reference page. 
• Prötzel, D., Vences, M., , Vieites, D. & Glaw, F. 2017. Splitting and Lumping: An Integrative Taxonomic Assessment of Malagasy Chameleons in the Calumma guibei complex results in the New Species C. gehringi sp. nov. Vertebrate Zoology 67(2): 231–249. PDF Reference page. 
• , Vences, M., Borrell, J., Ball, L., Nomenjanahary, D.H., Parker, D., Rakotondratsima, M., Razafimandimby, E., Starnes, T., Rabearivony, J. & Glaw, F. 2017. A new frog species of the subgenus Asperomantis (Anura, Mantellidae, Gephyromantis) from the Bealanana District of northern Madagascar. Zoosystematics and Evolution 93(2): 451–466. DOI: 10.3897/zse.93.14906 Reference page. 
• , Razafindraibe, J.H., Rakotoarison, A., Dixit, N.M., Bletz, M.C., Glaw, F. & Vences, M. 2017. Yet another small brown frog from high altitude on the Marojejy Massif, northeastern Madagascar (Anura: Mantellidae). Zootaxa 4347(3): 572–582. DOI: 10.11646/zootaxa.4347.3.9 Reference page. 
• , Vences, M., Rakotoarison, A., Anreone, F., Köhler, J., Glaw, F. & Crottini, A. 2017. Lumping or splitting in the Cophylinae (Anura: Microhylidae) and the need for a parsimony of taxonomic changes: a response to Peloso et al. (2017). Salamandra 53(3): 479–483. PDF Reference page. 

• Bellati, A., , Megson, S., Roberts, S.H., Andreone, F., Rosa, G.M., Noël, J., Randrianirina, J.E., Fasola, M., Glaw, F. & Crottini, A. 2018. Resurrection and re-description of Plethodontohyla laevis (Boettger, 1913) and transfer of Rhombophryne alluaudi (Mocquard, 1901) to the genus Plethodontohyla (Amphibia, Microhylidae, Cophylinae). Zoosystematics and Evolution 94(1): 109–135. DOI: 10.3897/zse.94.14698 Reference page. 
• Bletz, M.C., , Rakotoarison, A., Lehtinen, R.M., Glaw, F. & Vences, M. 2018. Stumbling upon a New Frog Species of Guibemantis (Anura: Mantellidae) on Top of the Marojejy Massif in Northern Madagascar. Copeia 106(2): 255–263. DOI: 10.1643/CH-17-655 Reference page. 
• Glaw, F., , Prötzel, D. & Vences, M. 2018. Eye and webbing colouration as predictors of specific distinctness: A genetically isolated new treefrog species of the Boophis albilabris group from the Masoala peninsula, northeastern Madagascar. Salamandra 54(3): 163–177. PDF Reference page. 
• Hawlitschek, O., , Ruthensteiner, B., Crottini, A. & Glaw, F. 2018. Computational molecular species delimitation and taxonomic revision of the gecko genus Ebenavia Boettger, 1878. The Science of Nature 105: 49. DOI: 10.1007/s00114-018-1574-9 Reference page. 
• Lehtinen, R.M., Glaw, F., Vences, M., Rakotoarison, A. & 2018. Two new Pandanus frogs (Guibemantis: Mantellidae: Anura) from northern Madagascar. European Journal of Taxonomy 451: 1–20. DOI: 10.5852/ejt.2018.451 Reference page. 
• , Hawlitschek, O., Razafindraibe, J.H., Megson, S., Ratsoavina, F.M., Rakotoarison, A., Bletz, M.C., Glaw, F. & Vences, M. 2018. A distinctive new frog species (Anura, Mantellidae) supports the biogeographic linkage of two montane rainforest massifs in northern Madagascar. Zoosystematics and Evolution 94(2): 247–261. DOI: 10.3897/zse.94.21037 Reference page. 
• , Rakotoarison, A., Ratsoavina, F.M., Hawlitschek, O., Vences, M. & Glaw, F. 2018. Two new Madagascan frog species of the Gephyromantis (Duboimantis) tandroka complex from northern Madagascar. Alytes 36(1–4): 130–158. Reference page. 
• Prötzel, D., Vences, M., Hawlitschek, O., , Ratsoavina, F.M. & Glaw, F. 2018. Endangered beauties: micro-CT cranial osteology, molecular genetics and external morphology reveal three new species of chameleons in the Calumma boettgeri complex (Squamata: Chamaeleonidae). Zoological Journal of the Linnean Society 184(2): 471–498. DOI: 10.1093/zoolinnean/zlx112 Reference page. 
• Prötzel, D., Lambert, S.M., Andrianasolo, G.T., Hutter, C.R., Cobb, K.A., & Glaw, F. 2018. The smallest ‘true chameleon’ from Madagascar: a new, distinctly colored species of the Calumma boettgeri complex (Squamata, Chamaeleonidae). Zoosystematics and Evolution 94(2): 409–423. DOI: 10.3897/zse.94.27305 Reference page. 
• Sentís, M., Chang, Y-Y., , Prötzel, D. & Glaw, F. 2018. Rising from the ashes: resurrection of the Malagasy chameleons Furcifer monoceras and F. voeltzkowi (Squamata: Chamaeleonidae), based on micro-CT scans and external morphology. Zootaxa 4483(3): 549–566. DOI: 10.11646/zootaxa.4483.3.7 Reference page. 

• Köhler, J., Vences, M., & Glaw, F. 2019. A new species of nocturnal gecko, genus Paroedura, from the karstic Tsingy de Bemaraha formation in western Madagascar. Salamandra 55(2): 73–81. PDF Reference page. 
• Rakotoarison, A., , Bletz, M.C., Razafindraibe, J.H., Glaw, F. & Vences, M. 2019. Description of the lucky Cophyla (Microhylidae, Cophylinae), a new arboreal frog from Marojejy National Park in north-eastern Madagascar. Zootaxa 4651(2): 271–288. DOI: 10.11646/zootaxa.4651.2.4 Reference page. 
• Ratsoavina, F.M., Raselimanana, A.P., , Rakotoarison, A., Razafindraibe, J.H., Glaw, F. & Vences, M. 2019. Finaritra! A new leaf-tailed gecko (Uroplatus) species from Marojejy National Park in north-eastern Madagascar. Zootaxa 4545(4): 563–577. DOI: 10.11646/zootaxa.4545.4.7 Reference page. 
• Ratsoavina, F.M., , Tolley, K.A., Raselimanana, A.P., Glaw, F. & Vences, M. 2019. A new species of Uroplatus (Gekkonidae) from Ankarana National Park, Madagascar, of remarkably high genetic divergence. Zootaxa 4683(1): 84–96. DOI: 10.11646/zootaxa.4683.1.4 Reference page. 
• , Köhler, J., Rakotoarison, A., Glaw, F. & Vences, M. 2019. A new dwarf chameleon, genus Brookesia, from the Marojejy massif in northern Madagascar. Zoosystematics and Evolution 95(1): 95–106. DOI: 10.3897/zse.95.32818 Reference page. 
• , Köhler, J., Vences, M. & Glaw, F. 2019. A new yellow-toed Platypelis species (Anura, Microhylidae, Cophylinae) from the Maroantsetra region, northeastern Madagascar. Evolutionary Systematics 3: 33417. DOI: 10.3897/evolsyst.3.33417 Reference page. 
• , Hutter, C.R., Rakotoarison, A., Riemann, J.C., Rödel, M.-O., Ndriantsoa, S.H., Glos, J., Roberts, S.H., Crottini, A., Vences, M. & Glaw, F. 2019. Morphological and ecological convergence at the lower size limit for vertebrates highlighted by five new miniaturised microhylid frog species from three different Madagascan genera. PLoS One 14(3): e0213314. DOI: 10.1371/journal.pone.0213314 Reference page. 
• , Glaw, F., Hutter, C.R., Bletz, M.C., Rakotoarison, A., Köhler, J. & Vences, M. 2019. Species complexes and the importance of Data Deficient classification in Red List assessments: The case of Hylobatrachus frogs. PLoS ONE 14(8): e0219437. DOI: 10.1371/journal.pone.0219437 Reference page. 

• Glaw, F., , Rakotoarison, A., Crottini, A., Raselimanana, A.P., Andreone, F., Köhler, J. & Vences, M. 2020. Genetic variability and partial integrative revision of Platypelis frogs (Microhylidae) with red flash marks from eastern Madagascar. Vertebrate Zoology 70(2): 141–156. DOI: 10.26049/VZ70-2-2020-04 Reference page. 
• Prötzel, D., , Ratsoavina, F.M., Vences, M. & Glaw, F. 2020. Untangling the trees: Revision of the Calumma nasutum complex (Squamata: Chamaeleonidae). Vertebrate Zoology 70(1): 23–59. DOI: 10.26049/VZ70-1-2020-3 Reference page. 
• Rakotoarison, A., , Köhler, J., Ratsoavina, F.M., Hawlitschek, O., Megson, S., Vences, M. & Glaw, F. 2020. Frogs of the genus Platypelis from the Sorata massif in northern Madagascar: description of a new species and reports of range extensions. Zoosystematics and Evolution 96(1): 263–274. DOI: 10.3897/zse.96.47088 Reference page. 
• Rancilhac, L., Bruy, T., , Pereira, E.A., Preick, M., Straube, N., Lyra, M.L., Ohler, A., Streicher, J.W., Andreone, F., Crottini, A., Hutter, C.R., Randrianantoandro, J.C., Rakotoarison, A., Glaw, F., Hofreiter, M. & Vences, M. 2020. Target-enriched DNA sequencing from historical type material enables a partial revision of the Madagascar giant stream frogs (genus Mantidactylus). Journal of Natural History 54(1-4): 87–118. DOI: 10.1080/00222933.2020.1748243 Reference page. 
• Ratsoavina, F.M., Glaw, F., Raselimanana, A.P., Rakotoarison, A., Vieites, D.R., Hawlitschek, O., Vences, M. & 2020. Towards completion of the species inventory of small-sized leaf-tailed geckos: two new species of Uroplatus from northern Madagascar. Zootaxa 4895(2): 251–271. DOI: 10.11646/zootaxa.4895.2.5 Reference page. 
• 2020. Diamond frogs forever: a new species of Rhombophryne Boettger, 1880 (Microhylidae, Cophylinae) from Montagne d’Ambre National Park, northern Madagascar. Zoosystematics and Evolution 96(2): 313–323. DOI: 10.3897/zse.96.51372 Reference page. 
• , Rasolonjatovo, S.M., Köhler, J., Rancilhac, L., Rakotoarison, A., Raselimanana, A.P., Ohler, A., Preick, M., Hofreiter, M., Glaw, F. & Vences, M. 2020. ‘Barcode fishing’ for archival DNA from historical type material overcomes taxonomic hurdles, enabling the description of a new frog species. Scientific Reports 10: 19109. DOI: 10.1038/s41598-020-75431-9 Reference page. 

• Glaw, F., Köhler, J., Hawlitschek, O., Ratsoavina, F.M., Rakotoarison, A., & Vences, M. 2021. Extreme miniaturization of a new amniote vertebrate and insights into the evolution of genital size in chameleons. Scientific Reports 11(1): 2522. DOI: 10.1038/s41598-020-80955-1 Reference page. 
• Gorin, V.A., , Korost, D.V. & Poyarkov, N.A. 2021. Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97(1): 21–54. DOI: 10.3897/zse.97.57968 Reference page. 
• Gippner, S., Travers, S.L., , Colston, T.J., Lyra, M.L., Mohan, A.V., Multzsch, M., Nielsen, S.V., Rancilhac, L., Glaw, F., Bauer, A.M. & Vences, M. 2021. A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution 165: 107311. DOI: 10.1016/j.ympev.2021.107311 Reference page. 
• Hawlitschek, O., , Webster, K.C., Ineich, I. & Glaw, F. 2021. Morphological, osteological, and genetic data support a new species of Madatyphlops (Serpentes: Typhlopidae) endemic to Mayotte Island, Comoros Archipelago. The Anatomical Record 304(10): 2249–2263. DOI: 10.1002/ar.24589 Reference page. 
• Miralles A., Bruy, T., Crottini, A., Rakotoarison, A., Ratsoavina, F.M., , Schmidt, R., Köhler, J., Glaw, F. & Vences, M. 2021. Completing a taxonomic puzzle: integrative review of geckos of the Paroedura bastardi species complex (Squamata, Gekkonidae). Vertebrate Zoology 71: 27–48. DOI: 10.3897/vz.71.e59495 Reference page. 
• , Schmidt, L., Crottini, A., Miralles, A., Rakotoarison, A., Raselimanana, A.P., Köhler, J., Glaw, F. & Vences, M. 2021. Into the Chamber of Horrors: A proposal for the resolution of nomenclatural chaos in the Scaphiophryne calcarata complex (Anura: Microhylidae), with a new species-level phylogenetic hypothesis for Scaphiophryninae. Zootaxa 4938(1): 392–420. DOI: 10.11646/zootaxa.4938.1.2 Reference page. 
• Vences, M., Köhler, J., Andreone, F., Craul, A.-K., Crottini, A., Du Preez, L.H., Preick, M., Rancilhac, L., Rödel, M.-O., , Streicher, J.W., Hofreiter, M. & Glaw, F. 2021. Target-enriched DNA sequencing clarifies the identity of name-bearing types of the Gephyromantis plicifer complex and reveals a new species of mantellid frog from Madagascar. Spixiana 44(2): 175–202. PDF Reference page. 

• Belluardo, F., , Santos, B., Andreone, F., Antonelli, A., Glaw, F., Muñoz-Pajares, A.J., Randrianirina, J.E., Raselimanana, A.P., Vences, M. & Crottini, A. 2022. Molecular taxonomic identification and species level phylogeny of the narrow-mouthed frogs of the genus Rhombophryne (Anura: Microhylidae: Cophylinae) from Madagascar. Systematics and Biodiversity 20(1): 1–13. DOI: 10.1080/14772000.2022.2039320 Reference page. 
• Mullin, K.E., Rakotomanga, M.G., Dawson, J., Glaw, F., Rakotoarison, A., Orozco-terWengel, P. & 2022. An unexpected new red-bellied Stumpffia (Microhylidae) from forest fragments in central Madagascar highlights remaining cryptic diversity. ZooKeys 1104ː 1–28. DOI: 10.3897/zookeys.1104.82396 Reference page. 
• Rakotoarison, A., Glaw, F., Rasolonjatovo, S.M., Razafindraibe, J.H., Vences, M. & 2022. Discovery of frogs of the Stumpffia hara species group (Microhylidae, Cophylinae) on Montagne d’Ambre in northern Madagascar, with description of a new species. Evolutionary Systematics 6: 21–33. DOI: 10.3897/evolsyst.6.76382 Reference page. 
• , Crottini, A., Hutter, C.R., Hildenbrand, A., Andreone, F., Fulgence, T.R., Köhler, G., Ndriantsoa, S.H., Ohler, A., Preick, M., Rakotoarison, A., Rancilhac, L., Raselimanana, A.P., Riemann, J.C., Rödel, M.-O., Rosa, G.M., Streicher, J.W., Vieites, D.R., Köhler, J., Hofreiter, M., Glaw, F. & Vences, M. 2022. An inordinate fondness for inconspicuous brown frogs: integration of phylogenomics, archival DNA analysis, morphology, and bioacoustics yields 24 new taxa in the subgenus Brygoomantis (genus Mantidactylus) from Madagascar. Megataxa 7(2): 113–311. DOI: 10.11646/megataxa.7.2.1 Reference page. 
• Vences, M., Multzsch, M., Gippner, S., Miralles, A., Crottini, A., Gehring, P.-S., Rakotoarison, A., Ratsoavina, F.M., Glaw, F. & 2022. Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar’s rainforest. Zootaxa 5179(1): 1–61. DOI: 10.11646/zootaxa.5179.1.1 Reference page. 
• Vences, M., Köhler, J., Crottini, A., Hofreiter, M., Hutter, C.R., Du Preez, L.H., Preick, M., Rakotoarison, A., Rancilhac, L., Raselimanana, A.P., Rosa, G.M., & Glaw, F. 2022. An integrative taxonomic revision and redefinition of Gephyromantis (Laurentomantis) malagasius based on archival DNA analysis reveals four new mantellid frog species from Madagascar. Vertebrate Zoology 72: 271–309. DOI: 10.3897/vz.72.e78830 Reference page. 

• Glaw, F., Köhler, J., Ratsoavina, F.M., Raselimanana, A.P., Crottini, A., Gehring, P.-S., Böhme, W., & Vences, M. 2023. A new large-sized species of leaf-tailed gecko (Uroplatus) from northern Madagascar. Salamandra 59(3): 239–261. PDF Reference page. 
• Koppetsch, T., Pabijan, M., Hutter, C.R., Köhler, J., Gehring, P.-S., Rakotoarison, A., Ratsoavina, F.M., , Vieites, D.R., Glaw, F. & Vences, M. 2023. An initial molecular resolution of the mantellid frogs of the Guibemantis liber complex reveals three new species from northern Madagascar. Vertebrate Zoology 73: 397–432. DOI: 10.3897/vz.73.e94063 Reference page. 
• Vences, M., Hutter, C.R., Glaw, F., Rakotoarison, A., Raselimanana, A.P. & 2023. A new species of Pandanus-dwelling frog from northern Madagascar related to Guibemantis pulcher. Zootaxa 5306(1): 97–115. DOI: 10.11646/zootaxa.5306.1.4 Reference page. 
• Vences, M., Multzsch, M., Köhler, J., Crottini, A., Andreone, F., Rakotoarison, A., & Glaw, F. 2023. Integrative revision of the Blommersia wittei complex, with description of a new species of frog from western and north-western Madagascar. Zootaxa 5319(): 178–198. DOI: 10.11646/zootaxa.5319.. Reference page. 

• Rakotoarison, A., Vences, M., Andreone, F., Crottini, A., Glaw, F., & Raselimanana, A.P. 2024. A new species of colorful Platypelis (Amphibia: Microhylidae) from the Tsaratanàna and Bemanevika massifs in northern Madagascar. Zootaxa 5501(1): 171–180. DOI: 10.11646/zootaxa.5501.1.8 Reference page. 
• Vences, M., Köhler, J., Hutter, C.R., Preick, M., Petzold, A., Rakotoarison, A., Ratsoavina, F.M., Glaw, F. & 2024. Communicator whistles: A Trek through the taxonomy of the Boophis marojezensis complex reveals seven new, morphologically cryptic treefrogs from Madagascar (Amphibia: Anura: Mantellidae). Vertebrate Zoology 74: 643–681. DOI: 10.3897/vz.74.e121110 Reference page. 
• Vences, M., Köhler, J., , Hutter, C.R., Rabe Maheritafika, H.M., Rafanoharana, J.M., Raherinjatovo, F., Rakotoarison, A., Andreone, F., Raselimanana, A.P. & Glaw, F. 2024. Four new species of forest-dwelling mantellid frogs from Madagascar allied to Gephyromantis moseri. Spixiana 46(2): 297–319. PDF Reference page. 

• Petzold, A., Glaw, F., Mullin, K.E., Rakotoarison, A., Raselimanana, A.P., Crottini, A., Orozco-terWengel, P., Köhler, J., Prötzel, D., Vences, M., Hofreiter, M. & 2025. A preliminary assessment of the diversity in the frog genus Anilany (Microhylidae: Cophylinae) with description of a new species from western Madagascar. Salamandra 61(1): 70–94. PDF Reference page. 
• Vences, M., Miralles, A., Ineich, I., Rakotoarison, A., Glasenapp, C., , Köhler, J., Glaw, F. & Raselimanana, A.P. 2025. An updated survey of molecular diversity in Madagascar's velvet geckos, genus Blaesodactylus, with description of a new species from the island’s arid West. Zootaxa 5620(2): 230–254. DOI: 10.11646/zootaxa.5620.2.2 Reference page.