Piperales

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Magnoliids
Ordo:
Familiae: Aristolochiaceae – Piperaceae – Saururaceae

Piperales Bercht. & J.Presl, Prir. Rostlin 261 (1820).

• Type genus: Piper L., Sp. Pl. 1: 28 (1753)
  

Note: Helmstetter et al. (2025) prefer to segregate Hydnoraand Prosopanche in their own family, Hydnoraceae C. Agardh. and also to segregate Lactoris into Lactoridaceae Engler. This is a valid taxonomic opinion, as is leaving them in Aristolochiaceae. It is recommended that WS follow APG (2016) pending further evidence

• Aristolochiales Bercht. & J.Presl, Prir. Rostlin 270 (1820).
• Asarales Horan., Char. Ess. Fam.: 68 (1847).
• Hydnorales Takht. ex Reveal, Novon 2: 239 (1992).
• Lactoridales Takht. ex Reveal, Phytologia 74: 174 (1993).

• Berchtold, F.v. & Presl, J.S. 1820. O Přirozenosti Rostlin, aneb Rostlinář, obsahugjcj: gednánj o žiwobytj rostlinném pro sebe a z ohledu giných žiwotů podlé stawu nyněgssjho znánj; k rozssjženj přirodnictwj; w potaženj na užitečnost w rolnictwj, hospodářstwj, řemeslech, uměnj i obchodu a w wztahowánj obzwlásstnjm na lekařstwj. 322 pp. + 2 pp. errata, Praha [Prague]: K. W. Enders. (Czech) Google Books HathiTrust. Reference page. 

• Angiosperm Phylogeny Group. 2009. An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III. Botanical Journal of the Linnean Society 161(2): 105–121. DOI: 10.1111/j.1095-8339.2009.00996.x Reference page. 
• Angiosperm Phylogeny Group. 2016. An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV. Botanical Journal of the Linnean Society 181(1): 1–20. DOI: 10.1111/boj.12385 Reference page. 
• Helmstetter, A.J., Ezedin, Z., de Lírio, E.J., de Oliveira, S.M. Chatrou, L.M., Erkens, R.H.J., Larridon, I., Leempoel, K., Maurin, O., Shyamali, R., Zuntini, A.R., Baker, W.J. Thomas, L.P., Couvreur, L.P., Forest, F. & Sauquet, H. 2025. Toward a phylogenomic classification of magnoliids. American Journal of Botany 111: e16451.DOI: 10.1002/ajb2.16451 Reference page. 
• Massoni, J., Forest, F. & Sauquet, H.J.X. 2014. Increased sampling of both genes and taxa improves resolution of phylogenetic relationships within Magnoliidae, a large and early-diverging clade of angiosperms. Molecular Phylogenetics and Evolution 70: 84–93. DOI: 10.1016/j.ympev.2013.09.010 ResearchGate Reference page. 
• Naumann, J., Salomo, K., Der, J.P., Wafula, E.K.,Bolin, J.F., Maass, E., Frenzke, L., Samain, M.S., Neinhuis, C. & Wanke, S. 2013. Single-copy nuclear genes place haustorial Hydnoraceae within Piperales and reveal a Cretaceous origin of multiple parasitic angiosperm lineages. PLoS One 8(11), p.e79204. DOI: 10.1371/journal.pone.0079204 Full text PDF Reference page. 
• Reveal, J.L. 2010. A checklist of familial and suprafamilial names for extant vascular plants. Phytotaxa 6: 1–402. DOI: 10.11646/phytotaxa.6.1.1 Reference page. 

• Stevens, P.F. 2001 onwards. Angiosperm Phylogeny Website. Version 14, July 2017 [and more or less continuously updated since]. Online. Reference page. 
• Tropicos.org 2014. Piperales. Missouri Botanical Garden. Published online. Accessed: 25 Jan. 2014. Reference page.

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