Santiago R. Ron

Santiago R. Ron, Ecuadorian herpetologist. Specialized in Neotropical anurans.

• Curator of Amphibians and Professor, Museo de Zoología and Departamento de Ciencias Biológicas, Pontificia Universidad Católica del Ecuador, Quito, Ecuador.

(List may be incomplete)

• 75 taxon names authored by Santiago R. Ron

(List may be incomplete)

• Guayasamin, J.M., , Cisneros-Heredia, D.F., Lamar, W.W. & McCracken, S.F. 2006. A new species of frog of the Eleutherodactylus lacrimosus assemblage (Leptodactylidae) from the western Amazon basin, with comments on the utility of canopy surveys in lowland rainforest. Herpetologica 62(2): 191–202. JSTOR Reference page. 

• , Venegas, P.J., Toral, E., Read, M., Ortiz, D. & Manzano, A.L. 2012. Systematics of the Osteocephalus buckleyi species complex (Anura, Hylidae) from Ecuador and Peru. ZooKeys 229: 1–52. DOI: 10.3897/zookeys.229.3580 Reference page. 

• Almendáriz, A., Brito, J., Batallas, D. & 2014. Una especie nueva de rana arbórea del género Hyloscirtus (Amphibia: Anura: Hylidae) de la Cordillera del Cóndor. Papéis Avulsos de Zoologia 54(4): 33–49. DOI: 10.1590/0031-1049.2014.54.04 Reference page. 

• Fouquet, A., Orrico, V.G.D., Ernst, R., Blanc, M., Martinez, Q., Vacher, J-P., Rodrigues, M.T., Ouboter, P., Jairam, R. & 2015. A new Dendropsophus Fitzinger, 1843 (Anura: Hylidae) of the parviceps group from the lowlands of the Guiana Shield. Zootaxa 4052(1): 39–64. DOI: 10.11646/zootaxa.4052.1.2 Reference page. 
• Ortega-Andrade, H.M., Rojas-Soto, O.R., Valencia, J.H., Espinosa de los Monteros, A., Morrone, J.J., & Cannatella, D.C. 2015. Insights from Integrative Systematics Reveal Cryptic Diversity in Pristimantis Frogs (Anura: Craugastoridae) from the Upper Amazon Basin. PLoS ONE 10(11): e0143392. DOI: 10.1371/journal.pone.0143392 Reference page. 
• , Mueses-Cisneros, J.J., Gutiérrez-Cárdenas, P.D.A., Rojas-Rivera, A., Lynch, R.L., Rocha, C.F.D. & Galarza, G. 2015. Systematics of the endangered toad genus Andinophryne (Anura: Bufonidae): phylogenetic position and synonymy under the genus Rhaebo. Zootaxa 3947(3): 347–366. DOI: 10.11646/zootaxa.3947.3.3. Preview (PDF) Full article (PDF) Reference page. 
• Yánez-Muñoz, M.H., Reyes-Puig, C., Bejarano-Muñoz, P. & 2015: Another new species of frog Pristimantis (Anura: Terrarana) in the western foothills of the Pichincha Volcano, Ecuador ACI Avances en Ciencias e Ingenierías 7(2): 52–60. Reference page. 

• , Venegas, P.J., Ortega-Andrade, H.M., Gagliardi-Urrutia, G. & Salerno, P.E. 2016. Systematics of Ecnomiohyla tuberculosa with the description of a new species and comments on the taxonomy of Trachycephalus typhonius (Anura, Hylidae). ZooKeys 630: 115–154. DOI: 10.3897/zookeys.630.9298 Reference page. 
• Székely, P., Cogălniceanu, D., Székely, D., Paez, N. & 2016. A new species of Pristimantis from southern Ecuador (Anura, Craugastoridae). ZooKeys 606: 77–97. DOI: 10.3897/zookeys.606.9121. Reference page. 
• Navarrete, M.J., Venegas, P.J. & 2016. Two new species of frogs of the genus Pristimantis from Llanganates National Park in Ecuador with comments on the regional diversity of Ecuadorian Pristimantis (Anura, Craugastoridae). ZooKeys 593: 139–162. DOI: 10.3897/zookeys.593.8063 Reference page. 

• Almendáriz C., A., Brito M., J., Batallas R., D., Vaca-Guerrero, J. & 2017. Una especie nueva de rana del género Chiasmocleis (Microhylidae: Gastrophryninae) de la Cordillera del Cóndor, Ecuador (In Spanish). Papéis Avulsos de Zoologia 57(10): 119–136. DOI: 10.11606/0031-1049.2017.57.10 Reference page. 
• Brito M., J., Almendáriz C., A., Batallas R., D. & 2017. Nueva especie de rana bromelícola del género Pristimantis (Amphibia: Craugastoridae), meseta de la Cordillera del Cóndor, Ecuador (In Spanish). Papéis Avulsos de Zoologia 57(15): 177–195. DOI: 10.11606/0031-1049.2017.57.15 Reference page. 
• Caminer, M.A., Milá, B., Jansen, M., Fouquet, A., Venegas, P.J., Chávez, G., Lougheed, S.C. & 2017. Systematics of the Dendropsophus leucophyllatus species complex (Anura: Hylidae): Cryptic diversity and the description of two new species. PLoS ONE 12(3): e0171785. DOI: 10.1371/journal.pone.0171785 Reference page. 
• Caminer, M.A., Milá, B., Jansen, M., Fouquet, A., Venegas, P.J., Chávez, G., Lougheed, S.C. & 2017. Correction: Systematics of the Dendropsophus leucophyllatus species complex (Anura: Hylidae): Cryptic diversity and the description of two new species. PLoS ONE 12(4): e0176902. DOI: 10.1371/journal.pone.0176902 Reference page. 
• Ortega-Andrade, H.M., Rojas-Soto, O.R., Espinosa de los Monteros, A., Valencia, J.H., Read, M. & 2017. Revalidation of Pristimantis brevicrus (Anura, Craugastoridae) with taxonomic comments on a widespread Amazonian direct-developing frog. The Herpetological Journal 17(1): 81–97. PDF Reference page. 
• Valencia, J.H., Dueñas, M.R., Székely, P., Batallas R., D., Pulluquitín, F. & 2017. A new species of direct-developing frog of the genus Pristimantis (Anura: Terrarana: Craugastoridae) from Cordillera del Cóndor, Ecuador, with comments on threats to the anuran fauna of the region. Zootaxa 4353(3): 447–466. DOI: 10.11646/zootaxa.4353.3.3. Reference page. 

• Rivadeneira, C.D., Venegas, P.J. & 2018. Species limits within the widespread Amazonian treefrog Dendropsophus parviceps with descriptions of two new species (Anura, Hylidae). ZooKeys 726: 25–77. DOI: 10.3897/zookeys.726.13864 Reference page. 
• Rojas, R.R., Fouquet, A., de Carvalho, V.T., , Chaparro, J.C., Vogt, R.C., Ávila, R.W., Farias, I.P., Gordo, M. & Hrbek, T. 2018. Redescription of the Amazonian tiny tree toad Amazophrynella minuta (Melin, 1941) (Anura: Bufonidae) from its type locality. Zootaxa 4482(3): 511–526. DOI: 10.11646/zootaxa.4482.3.4 Reference page. 
• Rojas, R.R., Fouquet, A., , Hernández-Ruz, E.J., Melo-Sampaio, P.R., Chaperro, J.C., Vogt, R.C., Carvalho, V.T., Pinheiro, L.C., Ávila, R.W., Farias, I.P., Gordo, M. & Hrbek, T. 2018. A Pan-Amazonian species delimitation: high species diversity within the genus Amazophrynella (Anura: Bufonidae). PeerJ 6: e4941. DOI: 10.7717/peerj.4941 Reference page. 
• , Caminer, M.A., Varela-Jaramillo, A. & Almeida-Reinoso, D. 2018. A new treefrog from Cordillera del Cóndor with comments on the biogeographic affinity between Cordillera del Cóndor and the Guianan Tepuis (Anura, Hylidae, Hyloscirtus). Zookeys 809: 97–124. DOI: 10.3897/zookeys.809.25207 Reference page. 
• , Duellman, W.E., Caminer, M.A. & Pazmiño, D. 2018. Advertisement calls and DNA sequences reveal a new species of Scinax (Anura: Hylidae) on the Pacific lowlands of Ecuador. PLoS ONE 13(9): e0203169. DOI: 10.1371/journal.pone.0203169 Reference page. 

• Paez, N. & 2019. Systematics of Huicundomantis, a new subgenus of Pristimantis (Anura, Strabomantidae) with extraordinary cryptic diversity and eleven new species. ZooKeys 868: 001–112. DOI: 10.3897/zookeys.868.26766. Reference page. 
• Reyes-Puig, C., Reyes-Puig, J.P., Velarde-Garcéz, D.A., Dávalos, N., Mancero, E., Navarette, M., Yánez-Muñoz, M.H., Cisneros-Heredia, D.F. & 2019. A new species of terrestrial frog Pristimantis (Strabomantidae) from the upper basin of the Pastaza River, Ecuador. Zookeys 832: 113–133. DOI: 10.3897/zookeys.832.30874 Reference page. 
• Reyes-Puig, C., Bittencourt-Silva, G.B., Torres-Sánchez, M., Wilkinson, M., Streicher, J.W., Maddock, S.T., Kotharambath, R., Müller, H., Larrera, F.N.A., Amieda-Reinoso, D., & Cisneros-Heredia, D.F. 2019. Rediscovery of the Endangered Carchi Andean Toad, Rhaebo colomai (Hoogmoed, 1985), in Ecuador, with Comments on Its Conservation Status and Extinction Risk. Check List 15(3): 415–419. DOI: 10.15560/15.3.415 Reference page. 
• Sánchez-Nivicela, J.C., Urgiles, V.L., Navarrete, M.J., Yánez-Muñoz, M.H. & 2019. A bizarre new species of Lynchius (Amphibia, Anura, Strabomantidae) from the Andes of Ecuador and first report of Lynchius parkeri in Ecuador. Zootaxa 4567(1): 1–24. DOI: 10.11646/zootaxa.4567.1.1. Reference page. 

• Chasiluisa, V.D., Caminer, M.A., Varela-Jaramillo, A. & 2020. Description and phylogenetic relationships of a new species of treefrog of the Osteocephalus buckleyi species group (Anura: Hylidae). Neotropical Biodiversity 6: 21–36. DOI: 10.1080/23766808.2020.1729306 Reference page. 
• Ferrão, M., Lima, A.P., , Santos, S.P. & Hanken, J. 2020. New Species of Leaf-litter Toad of the Rhinella margaritifera Species Group (Anura: Bufonidae) from Amazonia. Copeia 108(4): 967–986. DOI: 10.1643/CH2020043 Reference page. 
• Réjaud, A., Rodrigues, M.T., Crawford, A.J., Castroviejo-Fisher, S., Jaramillo-Martínez, A.F., Chaparro, J.C., Glaw, F., Gagliardi-Urrutia, G., Moravec, J., De la Riva, I.J., Pérez-Peña, P.E., Lima, A.P., Werneck, F.P., Hrbek, T., , Ernst, R., Kok, P.J.R., Driskell, A., Chave, J. & Fouquet, A. 2020. Historical biogeography identifies a possible role of Miocene wetlands in the diversification of the Amazonian rocket frogs (Aromobatidae: Allobates). Journal of Biogeography 47(11): 2472–2482. DOI: 10.1111/jbi.13937 Reference page. 
• Reyes-Puig, C., Wake, D.B., Kotharambath, R., Streicher, J.W., Koch, C., Cisneros-Heredia, D.F., Yánez-Muñoz, M.H. & 2020. Two extremely rare new species of fossorial salamanders of the genus Oedipina (Plethodontidae) from northwestern Ecuador. PeerJ 8: e9934. DOI: 10.7717/peerj.9934 Reference page. 
• , Carrión, J., Caminer, M.A., Sagredo, Y., Navarrete, M.J., Ortega, J.A., Vaela-Jaramillo, A,, Maldonado-Castro, G.A. & Terán, C. 2020. Three new species of frogs of the genus Pristimantis (Anura, Strabomantidae) with a redefinition of the P. lacrimosus species group. ZooKeys, 993: 121–155. DOI: 10.3897/zookeys.993.53559 Reference page. 
• Sánchez-Nivicela, J.C., Peloso, P.L.V., Urgilés, V.L., Yánez-Muñoz, M.H., Sagredo, Y., Páez, N., 2020. Description and phylogenetic relationships of a new trans-Andean species of Elachistocleis Parker 1927 (Amphibia, Anura, Microhylidae). Zootaxa 4779(3): 323–340. DOI: 10.11646/zootaxa.4779.3.2 Reference page. 
• Sturaro, M.J., Costa, J.C.L., Maciel, A.O., Lima-Filho, G.R., Rojas-Runjaic, F.M.J., Mejía, D.P., & Peloso, P.L.V. 2020. Resolving the taxonomic puzzle of Boana cinerascens (Spix, 1824), with resurrection of Hyla granosa gracilis Melin, 1941 (Anura: Hylidae). Zootaxa 4750(1): 1–30. DOI: 10.11646/zootaxa.4750.1.1 Reference page. 
• Pirani, R.M., Peloso, P.L.V., Prado, J.R., Polo, E.M., Knowles, L.L., , Rodrigues, M.T., Sturaro, M.J. & Werneck, F.P. 2020. Diversification history of clown tree frogs in Neotropical rainforests (Anura, Hylidae, Dendropsophus leucophyllatus group). Molecular Phylogenetics and Evolution 150: 106877. DOI: 10.1016/j.ympev.2020.106877 Reference page. 
• Reyes-Puig, C., Yánez Muñoz, M.H., Ortega, J.A. & 2020. Relaciones filogenéticas del subgénero Hypodictyon (Anura: Strabomantidae: Pristimantis) con la descripción de tres especies nuevas de la región del Chocó. Revista Mexicana de Biodiversidad 91 (e913013): 1–38. DOI: 10.22201/ib.20078706e.2020.91.3013 Reference page. 

• Carrión-Olmedo, J.C. & 2021. A new cryptic species of the Pristimantis lacrimosus group (Anura, Strabomantidae) from the eastern slopes of the Ecuadorian Andes. Evolutionary Systematics 5: 151–175. DOI: 10.3897/evolsyst.5.62661 Reference page. 
• Brito-Zapata, D., Reyes-Puig, C., Cisneros-Heredia, D.F., Zumel, D. & 2021. Description of a new minute frog of the genus Pristimantis (Anura: Strabomantidae) from Cordillera del Condor, Ecuador. Zootaxa 5072: 351–372. DOI: 10.11646/zootaxa.5072.4.3 Reference page. 
• Zumel, D., Buckley, D. & 2021. The Pristimantis trachyblepharis species group, a clade of miniaturized frogs: description of four new species and insights into the evolution of body size in the genus. Zoological Journal of the Linnean Society: 1–40. DOI: 10.1093/zoolinnean/zlab044 Reference page. 

• Bejarano-Muñoz, P., , Navarrete, M.J. & Yánez-Muñoz, M.H. 2022. Dos nuevas especies del grupo Pristimantis boulengeri (Anura: Strabomantidae) de la cuenca alta del río Napo, Ecuador. Cuadernos de Herpetología 36: 125–154 DOI: 10.31017/CdH.2022.(2020-103) Reference page. 
• Ortega, J.A., Brito-M., J. & 2022. Six new species of Pristimantis (Anura: Strabomantidae) from Llanganates National Park and Sangay National Park in Amazonian cloud forests of Ecuador. PeerJ 10 (e13761): 1–66. DOI: 10.7717/peerj.13761 Reference page. 

• Cisneros-Heredia, D.F., Yánez-Muñoz, M.H., Sánchez-Nivicela, J.C. & 2023. Two new syntopic species of glassfrogs (Amphibia, Centrolenidae, Centrolene) from the southwestern Andes of Ecuador. PeerJ 11: e15195. DOI: 10.7717/peerj.15195 Reference page. 
• Venegas, P.J., García-Ayachi, L.A., Toral, E., Malqui, J. & 2023. A new species of Spiny-backed Tree Frog, genus Osteocephalus (Anura, Hylidae), from the Yanachaga Chemillén National Park in central Peru. Evolutionary Systematics 7: 237–251. DOI: 10.3897/evolsyst.7.102360 Reference page. 

• Carrión-Bonilla, C.A., & Cook, J.A. 2024. Species richness in Myotis (Chiroptera: Vespertilionidae): species delimitation and expanded geographic sampling reveal high Neotropical diversity. Journal of Mammalogy 105(2): 241–258. DOI: 10.1093/jmammal/gyad124 Reference page. 
• López-Hervas, K., Santos, J.C., , Betancourth-Cundar, M., Cannatella, D.C. & Tarvin, R.D. 2024. Deep divergences among inconspicuously colored clades of Epipedobates poison frogs . Molecular Phylogenetics and Evolution 195: 108065. DOI: 10.1016/j.ympev.2024.108065 Reference page. 
• Menéndez-Guerrero, P.A., Santos, S.P.L., Salazar-Nicholls, M.J., Green, D.M. & 2024. Cryptic diversity in toads of the Rhinella marina species group (Anura, Bufonidae) with a subjectively beautiful new species from Western Ecuador. Zoological Journal of the Linnean Society 202(3): zlad197. DOI: 10.1093/zoolinnean/zlad197 Reference page. 
• , García, D., Brito-Zapata, D., Reyes-Puig, C., Figueroa-Coronel, E. & Cisneros-Heredia, D.F. 2024. A new glassfrog of the genus Centrolene (Amphibia, Centrolenidae) from the Subandean Kutukú Cordillera, eastern Ecuador. Zoosystematics and Evolution 100(3): 923–939. DOI: 10.3897/zse.100.116350 Reference page. 

• Aguirre, P.D., Apunte, K. & 2025. Systematics of the Dendropsophus leucophyllatus species group (Anura, Hylidae) from the Chocó region of Ecuador, with description of a new species. Evolutionary Systematics 9(1): 7-31. DOI: 10.3897/evolsyst.9.135431 Reference page. 
• Varela-Jaramillo, A., Streicher, J.W., Venegas, P.J. & 2025. Three new species of torrent treefrogs (Anura, Hylidae) of the Hyloscirtus bogotensis group from the eastern Andean slopes and the biogeographic history of the genus. ZooKeys 1231: 233–292. DOI: 10.3897/zookeys.1231.124926 Reference page. 
• Ortega, J.A., Cisneros-Heredia, D.F., Camper, J.D., Romero-Carvajal, A., Negrete, L. & 2025. Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography. Zoological Journal of the Linnean Society 203(1): 1–60. DOI: 10.1093/zoolinnean/zlae162 Reference page. 
• Moreno-Cárdenas, P., Pacheco, V., Ruelas, D. & , 2025. A new species of Cryptotis (Eulipotyphla: Soricidae) and introduction to the systematics of the Ecuadorian and Peruvian Cryptotis species. Mammalia (in press). DOI: 10.1515/mammalia-2024-0154 Reference page.