Stephen John Richards

Stephen John Richards (Richards), Australian entomologist and herpetologist.

• Terrestrial Vertebrates, South Australian Museum, North Terrace, Adelaide, South Australia 5000
• Department of Plant, Soil & Insect Sciences, University of Massachusetts, Amherst, Massachusetts, United States.

(List may be incomplete)

• 166 taxon names authored by Stephen John Richards

(List may be incomplete)

• , Dennis, A.J., Trenerry, M.P. & Werren, G.L. 1994. A new species of Cophixalus (Anura: Microhylidae) from northern Queensland. Memoirs of the Queensland Museum 37: 307–310. BHL Reference page. 

• , Burton, T.C., Cunningham, M.J. & Dennis, A.J. 1995. A new species of Callulops from New Guinea and comments on the status of C. humicola computs (Zweifel) (Anura: Microhylidae: Asterophryinae). Transactions of the Royal Society of South Australia 119: 157–162. Archive Reference page. 

• Günther, R., , & Iskandar D.T. 2001. Two new species of the genus Oreophryne from Irian Jaya, Indonesia (Amphibia, Anura, Microhylidae). Spixiana 24(3): 257–274. ResearchGate Reference page. 
• & Iskandar, D.T. 2001. A new species of tree frog (Anura, Hylidae, Litoria) from the mountains of Irian Jaya, Indonesia. Alytes 18(3–4): 141–152. PDFReference page. 

• Hiaso, J. & 2006. A new species of Litoria (Anura: Hylidae) from Tagula Island, Papua New Guinea. Science in New Guinea. Port Moresby 28: 34–38. Reference page. 
• , Oliver, P.M., Dahl, C. & Tjaturadi, B. 2006. A new species of large green treefrog (Anura: Hylidae: Litoria) from northern New Guinea. Zootaxa 1208: 57–68. Abstract & excerpt. .ResearchGate Reference page. 
• & Iskandar, D.T. 2006. A New Species of Torrent-Dwelling Frog (Hylidae, Litoria) from the Mountains of New Guinea. Current Herpetology 25(2): 57–63. DOI: 10.3105/1345-5834(2006)25[57:ANSOTF]2.0.CO;2 PDF ResearchGate Reference page. 

• , Dahl, C.S. & Hiaso, J. 2007. Another new species of Choerophryne (Anura: Microhylidae) from Southern Highlands Province, Papua New Guinea. Transactions of the Royal Society of South Australia 131: 135–141. Reference page. 
• Rösler, H., & Günther, R. 2007. Bemerkungen zur Morphologie und Taxonomie der östlich der Wallacea vorkommenden Geckos der Gattung Cyrtodactylus Gray, 1827, mit Beschreibungen von zwei neuen Arten (Reptilia: Sauria: Gekkonidae). Salamandra 43(4): 193–230. PDF Reference page. 
• Oliver, P.M., , Tjaturadi, B., & Iskandar D.T. 2007. A New Large Green Species Of Litoria (Anura: Hylidae) From Western New Guinea. Zootaxa 1519: 17–26. Abstract & excerpt. ResearchGate PDF. Reference page. 

• Menzies, J.I., & Tyler, M.J. 2008. Systematics of the Australo-Papuan tree frogs known as Litoria bicolor (Anura: Hylidae) in the Papuan region. Australian Journal of Zoology 56(4): 257–280. DOI: 10.1071/ZO07068 Reference page. 
• Oliver, P., Tjaturadi, B., Mumpuni, Krey, K. & 2008. A new species of large Cyrtodactylus (Squamata: Gekkonidae) from Melanesia. Zootaxa 1894(1): 59–68. DOI: 10.11646/zootaxa.1894.1.5 ; ResearchGate Reference page. 
• Polhemus, D.A., Michalski, J. & 2008. Pseudagrion fumipennis, a remarkable new species of damselfly from New Guinea (Odonata: Zygoptera: Coenagrionidae). Tijdschrift voor Entomologie 151(1): 51–56. DOI: 10.1163/22119434-900000249 Reference page. 

• Günther, R., , Tjaturadi, B., & Iskandar D.T. 2009. A new species of the microhylid frog genus Oreophryne from the Mamberamo Basin of northern Papua Province, Indonesian New Guinea. Vertebrate Zoology 59(2): 147-155. PDF. Reference page. 
• , Oliver, P.M., Krey & Tjaturadi, B. 2009. A new species of Litoria (Amphibia: Anura: Hylidae) from the foothills of the Foja Mountains, Papua Province, Indonesia. Zootaxa 2277: 1–13. Abstract & excerpt. . Reference page. 

• Austin, C.C., Rittmeyer, R.N., Richards, S.J. & Zug, G.R. 2010. Phylogeny, historical biogeography and body size evolution in Pacific Island Crocodile skinks Tribolonotus (Squamata; Scincidae). Molecular Phylogenetics and Evolution 57(1): 227–236. DOI: 10.1016/j.ympev.2010.06.005 ; hdl: 10088/16716 . Reference page. 

• Oliver, P.M., Krey K., Mumpuni & Richards, S.J. 2011. A new species of bent-toed gecko (Cyrtodactylus, Gekkonidae) from the North Papuan Mountains. Zootaxa 2930: 22–32. Preview ResearchGate . Reference page. 

• Günther, R. & Richards, S. 2012 "2011". Description of two new species of the microhylid frog genus Oreophryne (Amphibia: Anura: Microhylidae) from southern Papua New Guinea. Vertebrate Zoology 61(3): 343–372. PDF Reference page. 
• Günther, R., Richards, S.J., Bickford, D.P. & Johnston, G.R. 2012. A new egg-guarding species of Oreophryne (Amphibia, Anura, Microhylidae) from southern Papua New Guinea. Zoosystematics and Evolution 88(2): 223–230. DOI: 10.1002/zoos.201200019 Reference page. 
• Michalski, J., & Theischinger, G. 2012. An interesting new species of Nososticta Hagen from southern New Guinea (Odonata, Disparoneuridae). Zootaxa 3590: 73–78. Preview Reference page. 
• Oliver, P.M., , & Tjaturadi, B., 2012. Two new species of Callulops (Anura: Microhylidae) from montane forests in New Guinea. Zootaxa 3178: 33–44. Preview. Reference page. 
• Orr, A.G., Kalkman, V.J. & 2012. A review of the New Guinean genus Paramecocnemis Lieftinck (Odonata: Platycnemididae), with the description of three new species. The Australian Entomologist 39(3): 161–177. Full issue (PDF) Reference page. 

• Günther, R., Richards, S.J. & Dahl, C.S. 2014. Nine new species of microhylid frogs from the Muller Range in western Papua New Guinea (Anura, Microhylidae). Vertebrate Zoology 64(1): 59–94. Full article (PDF) Reference page. 
• Orr, A.G. & 2014. Palaiargia traunae sp. n. (Odonata: Platycnemididae), a new idiocnemidine damselfly from Papua New Guinea. Australian entomologist 41(3): 153–159. Full article (PDF) on informit Reference page. 
• Orr, A.G., Kalkman, V.J. & 2014. Four new species of Palaiargia Förster, 1903 (Odonata: Platycnemididae) from New Guinea with revised distribution records for the genus. International Journal of Odonatology 16(4) (2013): 309–325. DOI: 10.1080/13887890.2013.855949 . Reference page. 
• Richards, S.J., Oliver, P. & Brown, R.B. 2014. A new scansorial species of Platymantis Günther 1858 (Anura: Ceratobatrachidae) from Manus Island, Admiralty Archipelago, Papua New Guinea. pp. 123–134 in Telnov, D. (ed.). Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea. Volume II. The Entomological Society of Latvia: Lettland, Latvia. ISBN 9789984976877 Reference page. 

• Theischinger, G., Gassmann, D. & Richards, S.J. 2015. Microcnemis gracilis, a new genus and species of Idiocnemidinae (Zygoptera: Platycnemididae) from Papua New Guinea. Zootaxa 3990(3): 429–436. DOI: 10.11646/zootaxa.3990.3.7. Preview (PDF) Reference page. 
• Brown, R.M., Siler, D.S., , Diesmos, A.C. & Cannatella, D.C. 2015. Multilocus phylogeny and a new classification for Southeast Asian and Melanesian forest frogs (family Ceratobatrachidae). Zoological journal of the Linnean Society 174(1): 130–168. DOI: 10.1111/zoj.12232 Reference page. 
• Xu, G., Tang, W., Skelley, P.E., Liu, N. & Richards, S.J. 2015. Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis (Cycadaceae) in Asia. Zootaxa 3986(3): 251–278. DOI: 10.11646/zootaxa.3986.3.1. Preview (PDF) Reference page. 
• Iannella, A., Oliver, P.M. & Richards, S. 2015. Two new species of Choerophryne (Anura, Microhylidae) from the northern versant of Papua New Guinea’s central cordillera. Zootaxa 4058(3): 332–340. DOI: 10.11646/zootaxa.4058.3.2. Preview (PDF). Reference page. 
• Günther, R., Richards, S.J., Tjaturadi, B., Krey, K. 2015. Two new species of the genus Cophixalus from the Raja Ampat Islands west of New Guinea (Amphibia, Anura, Microhylidae). Zoosystematics and Evolution 91(2): 199-213. DOI: 10.3897/zse.91.5411 . Reference page. 

• Günther, R., , & Tjaturadi, B. 2016. A New Species of the Frog Genus Pseudocallulops from the Foja Mountains in Northwestern New Guinea (Amphibia, Microhylidae). Russian Journal of Herpetology 23(1): 63-69. Reference page. 
• Gassmann, D., & Polhemus, D.A. 2016. Idiocnemis schorri sp. nov., a new damselfly species from southern Papua New Guinea (Odonata: Platycnemididae). Zootaxa 4171(3): 491–504. DOI: 10.11646/zootaxa.4171.3.5 Reference page. 
• McDonald, K.R., Rowley, J.J.L., & Frankham, G.J. 2016. A new species of treefrog (Litoria) from Cape York Peninsula, Australia. Zootaxa 4171(1): 153–169. DOI: 10.11646/zootaxa.4171.1.6 Reference page. 
• Oliver, P.M., Richards, S., Mumpuni & Rösler, H. 2016. The Knight and the King: two new species of giant bent-toed gecko (Cyrtodactylus, Gekkonidae, Squamata) from northern New Guinea, with comments on endemism in the North Papuan Mountains. ZooKeys 562: 105–130. DOI: 10.3897/zookeys.562.6052 ResearchGate .Reference page. 
• Oliver, P.M., Clegg, J.R., Fisher, R.N., Richards, S.J., Taylor, P.N. & Jocque, M.M.T. 2016. A new biogeographically disjunct giant gecko (Gehyra: Gekkonidae: Reptilia) from the East Melanesian Islands. Zootaxa 4208(1): 61–76. DOI: 10.11646/zootaxa.4208.1.3. Full article (PDF). Reference page. 
• Orr, A.G. & 2016. Three new species of Papuagrion Ris, 1913 (Odonata: Coenagrionidae) from the Hindenburg Wall region of western Papua New Guinea. Zootaxa 4072(3): 319–332. DOI: 10.11646/zootaxa.4072.3.2 Reference page. 
• Günther, R. & Richards, S. 2016. Description of two new species of the microhylid frog genus Oreophryne (Amphibia: Anura: Microhylidae) from southern Papua New Guinea. Vertebrate Zoology 66(2): 157–168. Full article (PDF) Reference page. 
• Günther, R., 2016. Description of a striking new Mantophryne species (Amphibia, Anura, Microhylidae) from Woodlark Island, Papua New Guinea. Zoosystematics and Evolution. 92(1): 111-118. DOI: 10.3897/zse.92.7629 Reference page. 

• Günther, R. & Richards, S. 2017. Three new species of the microhylid frog genus Choerophryne (Amphibia, Anura, Microhylidae) from Papua New Guinea. Zoosystematics and Evolution 93(2): 265–279. DOI: 10.3897/zse.93.11576. Reference page. 

• Günther, R. & 2018. A new species of the microhylid frog genus Choerophryne from Papua New Guinea. Alytes 36: 159–169. Reference page. 
• Günther, R., , & Tjaturadi, B. 2018. Two new microhylid frog species of the genus Xenorhina Peters, 1863 from the Raja Ampat Islands, Indonesia. Vertebrate zoology 68(2): 109–122. DOI: 10.30906/1026-2296-2016-23-1-63-69 . Reference page. 
• O’Shea, M., Herlihy, B., Paivu, B., Parker, F., & Kaiser, H. 2018. Rediscovery of the rare Star Mountains Worm-eating Snake, Toxicocalamus ernstmayri O’Shea et al., 2015 (Serpentes: Elapidae: Hydrophiinae) with the description of its coloration in life. Amphibian & Reptile Conservation 12(1): 27–34. PDF Reference page. 
• Travers, S.L., Richards, S.J., Broadhead, T.S. & Brown, R.M. 2018. A new miniature Melanesian Forest Frog (Ceratobatrachidae: Cornufer) from New Britain Island, constituting the first record of the subgenus Batrachylodes from outside of the Solomon Archipelago. Zootaxa 4370(1): 23–44. DOI: 10.11646/zootaxa.4370.1.2. Reference page. 
• Ruane, S., , McVay, J.D., Tjaturadi, B., Krey, K. & Austin, C.C. 2018. Cryptic and non-cryptic diversity in New Guinea ground snakes of the genus Stegonotus Duméril, Bibron and Duméril, 1854: a description of four new species (Squamata: Colubridae). Journal of Natural History 52(13-16): 917–944. DOI: 10.1080/00222933.2017.1391959 PDF ResearchGate Reference page. 

• Gassmann, D. & Richards, S.J. 2019. Two new damselflies of the genus Idiocnemis Selys from Gulf Province, Papua New Guinea (Odonata: Platycnemididae). Zootaxa 4560(1): 121–140. DOI: 10.11646/zootaxa.4560.1.6 Reference page. 
• Oliver, P.M., Günther, R., Mumpuni & Richards, S.J. 2019. Systematics of New Guinea treefrogs (Litoria: Pelodryadidae) with erectile rostral spikes: an extended description of Litoria pronimia and a new species from the Foja Mountains. Zootaxa 4604(2): 335–348. DOI: 10.11646/zootaxa.4604.2.6 Reference page. 
• Oliver, P.M., Richards, S.J. & Donnellan, S.C. 2019. Two new species of treefrog (Pelodrydidae: Litoria) from southern New Guinea elucidated by DNA barcoding. Zootaxa 4609(3): 469–484. DOI: 10.11646/zootaxa.4609.3.4 Reference page. 
• Richards, S. & Günther, R. 2019 Three new scansorial species of microhylid frogs (Anura: Cophixalus, Oreophryne) from Papua New Guinea. Salamandra 55(2): 55–72. Full article (PDF). Reference page. 
• Phimmachak, S., Richards, S.J., Sivongxay, N., Seateun, S., Chuaynkern, Y., Makchai, S., Som, H.E. & Stuart, B.L. 2019. A new caruncle-bearing fanged frog (Limnonectes, Dicroglossidae) from Laos and Thailand. ZooKeys 846: 133–156. DOI: 10.3897/zookeys.846.33200A Reference page. 

• Karkkainen, D.T., , Kraus, F., Tjaturadi, B., Krey, K. & Oliver, P.M. 2020. A new species of small Lepidodactylus (Squamata: Gekkonidae) from Salawati Island, Indonesia. Israel Journal of Ecology and Evolution 66(3–4): 180–189. DOI: 10.1163/22244662-bja10001 Reference page. 
• Oliver, P.M., Hartman, R., Turner, C.D., Wilde, T.A., Austin, C.C. & Richards, S.J. 2020. A new species of Cyrtodactylus Gray (Gekkonidae: Squamata) from Manus Island, and extended description and range extension for Cyrtodactylus sermowaiensis (De Rooij). Zootaxa 4728(3): 341–356. DOI: 10.11646/zootaxa.4728.3.3 Reference page. 
• Tallowin, O.J.S., Meiri, S., Donnellan, S.C., , Austin, C.C. & Oliver, P.M. 2020. The other side of the Sahulian coin: biogeography and evolution of Melanesian forest dragons (Agamidae). Biological Journal of the Linnean Society 129(1): 99–113. DOI: 10.1093/biolinnean/blz125 Reference page. 
• Günther, R., , Tjaturadi, B., Krey, K. 2020. Two new microhylid frog species of the genus Xenorhina Peters, 1863 from the Raja Ampat Islands, Indonesia. Vertebrate zoology 70(3): 333–347. DOI: 10.26049/VZ70-3-2020-06 PDF. Reference page. 
• Günther, R. & 2020. Two new frog species of the genus Copiula Mehely, 1901 (Anura, Microhylidae, Asterophryinae) from southern Papua New Guinea. Russian Journal of Herpetology 27: 41–53. DOI: 10.30906/1026-2296-2020-27-1-41-53 Reference page. 
• & Donnellan, S.C. 2020. Litoria aplini sp. nov., a new species of treefrog (Pelodryadidae) from Papua New Guinea. Records of the Australian Museum 72(5): 325–337. DOI: 10.3853/j.2201-4349.72.2020.1729 Reference page. 

• Oliver, P., Günther, R., Tjaturadi, B. & Richards, S.J. 2021. A new species of large green treefrog (Litoria, Pelodryadidae) from Papua, Indonesia. Zootaxa 4903(1): 117–126. DOI: 10.11646/zootaxa.4903.1.7 Reference page. 
• , Tjaturadi, B., Krey, K. & Donnellan, S.C. 2021. A new stream-dwelling frog of the genus Litoria Tschudi, 1838 (Anura: Pelodryadidae) from Salawati Island, Indonesia. Pp. 419–426, pl. 83. In: Telnov, D., Barclay, M.V.L. & Pauwels, O.S.G. (eds.) Biodiversity, biogeography and nature conservation in Wallacea and New Guinea. Volume IV. The Entomological Society of Latvia, Riga. 448 pp. 19–33 pls. ISBN 978-9984-9768-9-1. . Reference page. 
• Günther, R. & 2021. Description of six new species of Xenorhina Peters, 1863 from southern Papua New Guinea (Amphibia, Anura, Microhylidae). Zoosystematics and Evolution 97: 355–382. DOI: 10.3897/zse.97.59696 Reference page. 
• Günther, R. & 2021. Three new tiny species of the microhylid frog genus Cophixalus Boettger, 1892 from central Papua New Guinea. Proceedings of the Biological Society of Washington 134(1): 339–362. DOI: 10.2988/0006-324X-134.339 Reference page. 
• Oliver, P.M., Karkkainen, D. & 2021. A new species of Cyrtodactylus (Squamata: Gekkonidae) from the northern versant of New Guinea’s Central Cordillera. Zootaxa 5057(2): 260–270. DOI: 10.11646/zootaxa.5057.2.6 Reference page. 
• Günther, R., Dahl, C.S. & 2021. Description of a new Xenorhina species (Anura, Microhylidae) from northwestern Papua New Guinea. Vertebrate Zoology. Senckenberg. 71: 621–630. DOI: 10.3897/vz.71.e66954 Reference page. 

• & Bickford, D.P. 2023. A New Species of Small Green Treefrog (Pelodryadidae: Litoria) from the Lakekamu Basin in Southern Papua New Guinea. Current Herpetology 42(1): 43–54. DOI: 10.5358/hsj.42.43 Reference page. 
• & Donnellan, S.C. 2023. Two new species of green treefrogs (Pelodryadidae: Litoria) from the northern slopes of Papua New Guinea’s Central Cordillera. Zootaxa 5271(3): 477–502. DOI: 10.11646/zootaxa.5271.3.3 Reference page. 
• Günther, R., Dahl, C. & 2023. Another giant species of the microhylid frog genus Cophixalus Boettger, 1892 from the mountains of Papua New Guinea and first records of procoracoids in the genus. Zoosystematics and Evolution 99(1): 173–183. DOI: 10.3897/zse.99.97006 Reference page. 
• Günther, R., Iskandar, D.T. & 2023. A new large Oreophryne species from the mountains of Papua Province, Indonesian New Guinea (Amphibia, Anura, Microhylidae). Vertebrate Zoology 73: 153–159. DOI: 10.3897/vz.73.e94207 Reference page. 
• , Donnellan, S.C. & Oliver, P.M. 2023. Five new species of the pelodryadid genus Litoria Tschudi from the southern versant of Papua New Guinea’s Central Cordillera, with observations on the diversification of reproductive strategies in Melanesian treefrogs. Zootaxa 5263(2): 151–190. DOI: 10.11646/zootaxa.5263.2.1 Reference page. 
• Donnellan, S.C., Catullo, R.A., Rowley, J.J.L., Doughty, P., Price, L.C., Hines, H.B. & 2023. Revision of Litoria rothii (Anura: Pelodryadidae) from northern Australia. Zootaxa 5352(1): 73–108. DOI: 10.11646/zootaxa.5352.1.3 Reference page. 

• Oliver, P.M., Boothroyd, N., Tjaturadi, B., Riyanto, A., Iskandar, D.T. & 2024. A new species of narrow-banded Cyrtodactylus (Gekkonidae) from northern New Guinea. Zootaxa 5506(1): 79–92. DOI: 10.11646/zootaxa.5506.1.4 Reference page. 
• Slavenko, A., , Donnellan, S.C., Allison, A. & Oliver, P.M. 2024. Gold in the mountains: Striking new species of Papuascincus (Sphenomorphini: Scincidae) from New Guinea. Vertebrate Zoology 74: 133–149. DOI: 10.3897/vz.74.e112782 Reference page. 
• & Oliver, P.M. 2024. Two new species of torrent-breeding treefrogs (Anura: Pelodryadidae: Litoria) from hill forests on the southern edge of New Guinea’s Central Cordillera. Vertebrate Zoology 74: 417–433. DOI: 10.3897/vz.74.e123251 Reference page. 

• Oliver, P.M. & 2025. A new species of Lepidodactylus (Gekkonidae) from Misima Island, Papua New Guinea. Zootaxa 5601(2): 368–376. DOI: 10.11646/zootaxa.5601.2.10 Reference page. 
• Slavenko, A., Shea, G.M., & Oliver, P.M. 2025. A new species of montane Prasinohaema (Sphenomorphini: Scincidae) from the South-fold Mountains of New Guinea. Zootaxa 5621(2): 211–230. DOI: 10.11646/zootaxa.5621.2.3. Reference page. 
• Günther, R., Nagombi, E. & 2025. Two new species of microhylid frogs (Cophixalus, Oreophryne) from the Purari River Basin, Papua New Guinea. Zootaxa 5604(3): 234–254. DOI: 10.11646/zootaxa.5604.3.2 Reference page. 
• , Johnston, G.R. & Oliver, P.M. 2025. A new montane species of stream-dwelling treefrog (Pelodryadidae) from the Star Mountains, Papua New Guinea. Zootaxa 5661(2): 221–236. DOI: 10.11646/zootaxa.5661.2.4 Reference page.