Thomas Pain

Thomas Pain, (13.VII.1915 – 2.VI.2003) British conchologist and malacologist. Described 49 species of mollusks.

• 1946. Two new species of Pila (=Ampullaria) from South America. Proceedings of the Malacological Society of London 26: 180–181, pl. 6.
• 1946. On Pila canaliculata and its locality. Proceedings of the Malacological Society of London 27: 58–59.

• & Blok, A. 1948. Shell collection of the late Revd. E.G. Alderson with special reference to the genus Pila Röding. Journal of conchology 22: 299–302.

• 1949. Placostylus falcicola (Gassies). Journal of conchology 23: 39–40.
• 1949. The distribution of Pila speciosa (Philippi). Journal of conchology 23: 69–70.
• 1949. Three new species of Pomacea. Proceedings of the Malacological Society of London 27: 257–258, pl. 13.
• 1949. On the types of three species of Pomacea described by G.B. Sowerby III. Proceedings of the Malacological Society of London 28: 39–40, pl. 1: 2.

• 1950. A new species of Pomacea (Limnopomus) from Venezuela. Journal of conchology 23: 109–111.
• 1950. Pomacea (Ampullariidae) of British Guiana. Proceedings of the Malacological Society of London 28: 63–74, pl. 6–8.

• 1951. Pomacea hanleyana (Alderson). Journal of conchology 23: 145–146, fig1-9.
• 1951. The type of Pila nyanzae (Smith) Proceedings of the Malacological Society of London 28: 230–231, pl. 28.

• & Blume, W. 1952. A new species of Pomacea from Bolivia. Journal of conchology 23: 267–268, 1: 7.
• 1952. Notes on the Pomacea of Surinam with special reference to Ampullaria sowerbyi Vernhout. Basteria 16: 30–32 (1952).
• 1952. Notes on Pila ovata Oliver (sic) and its distribution in Africa. Revue de zoologie et de botanique africaines 46: 286–292.

• 1953. A giant E. African Achatina. Journal of conchology 23: 298.
• 1953. Sinistrality in Achatina fulica Bowdich. Journal of conchology 23: 370–371, 1: 14.
• 1953. Pomacea ghiesbreghti (Reeve) in Guatemala. Proceedings of the Malacological Society of London 29: 222–223.

• 1954. New freshwater Gastropod Mollusks of the African genus Lanistes. Breviora 31: 1–4, fig. 1–2.
• 1954. Notes on a collection of non-marine Mollusca from semi-arid areas of East Africa. Journal of conchology 23: 398–401.

• 1955. Notes on some New Caledonia Placostylus. Journal de conchyliologie 95: 11–19, 9 figs.

• 1956. Revision of the Melaniidae of British Guiana and Surinam. Basteria 20: 91–105, fig1-11.
• 1956. On a collection of Pomacea from Colombia with description of a new subspecies. Journal of conchology 24: 71–79.
• 1956. Notes on the generic names Pomacea and Ampullarius. Journal of conchology 24: 79.
• 1956. Further notes of the non-marine Mollusca from semi-arid areas of East Africa. Journal of conchology 24: 142–143.

• 1957. Pomacea of the Sierra de Merida, Venezuela. Journal of conchology 24: 175–176.
• 1957. On the occurrence of Achatina stuhlmanni v. Martens in Kenya. Revue de zoologie et de botanique africaines 56: 345–348.

• & Arias, C. 1958. Descripción de una especie nueva de Pomacea de Venezuela (Mesogastropoda, Architaenioglossa, Mollusca). Novedades científicas 24: 5–11, 1. i, ii.
• & Crowley, T.E. 1958. On an unfigured species of Plekocheilus (Eurytus) from Colombia. Journal of conchology 24: 234–235, pl 7. Reference page. 
• & Willson, J.A. 1958. Malayan sand snails (Melanoides tuberculata (O.F. Müller)). Fishkeeping & Water Life 13: 117–118.
• & Willson, J.A. 1958. Apple snails (Ampullaria). Fishkeeping & Water Life 13: 549–550.
• 1958. On a new subspecies of Placostylus fibratus (Martyn) from New Caledonia. Journal of conchology 24: 276–277, pl. 8.

• & Crowley, T.E. 1959. A monographic revision of the African land snails of the genus Burtoa Bourguignat. Annales du Musée Royal du Congo Belge, N. Sér. in-8°, Sciences Zoologiques 79: 1–35, pl. 1–3. Reference page. 
• & Willson, J.A. 1959. River snails. Aquarist and Pondkeeper 24: 130–131, 4 figs.
• 1959. A giant Achatina from the Belgian Congo (Achatina (Achatina) balteata infrafusca von Martens). Journal of conchology 24: 356.
• 1959. Orthalicus (Metorthalicus) labeo (Broderip) a rare Peruvian land snail. Journal of conchology 24: 357–358.

• 1960. On the distribution and synonymy of Strophocheilus (Megalobulimus) lichtensteini (Albers). Journal of conchology 24: 378.
• 1960. Pomacea (Ampullariidae) of the Amazon river system. Journal of conchology 24: 421–432.

• & Crowley, T.E. 1961. Pfeiffer's unfigured species of Strophocheilus (Megalobulimus). Breviora 138: 1–8, 6 figs. Reference page. 
• & Crowley, T.E. 1961. On the occurrence of Burtoa nilotica (Pfeiffer) in Ethiopia. Journal of conchology 25: 38. Reference page. 
• & Crowley, T.E. 1961. Description of new Achatinidae from the Congo and Nyasaland (Mollusca, Pulmonata). Revue de zoologie et de botanique africaines 64: 138–152, fig1-8. Reference page. 
• & Crowley, T.E. 1961. A monograph of the African land snails of the genus Limicolariopsis d'Ailly (Mollusca, Achatinidae) Annales du Musée Royal de l'Afrique Centrale: Serie in 8° Sciences Zoologiques 101: 1–36, pl. 1–3. Reference page. 
• & Woodward, F.R. 1961. A revision of the freshwater mussels of the family Etheriidae. Journal of conchology 25: 2–8.
• 1961. Revision of the African Ampullariidae species of the genus Pila Röding 1798 (Mesogastropoda, Architaenioglossa, Mollusca). Annales du Musée Royal de l'Afrique Centrale: Serie in 8° Sciences Zoologiques 96: 1–27.
• 1961. Introduction to Molluscan taxonomy. The Conchologists' Newsletter 2: 9–10.
• 1961. Introduction to Molluscan taxonomy. The Conchologists' Newsletter 3: 14–15.

• & Verdcourt, B. 1962. On the occurrence of Achatina tavaresiana Morelet in Tanganyika Territory. Journal of conchology 25: 82. Reference page. 
• & Woodward, F.R. 1962. The African freshwater bivalve Aspatharia (Spathopsis) rubens (Lamarck) its synonymy and distribution. Journal of conchology 25: 73–78 pl. 5.
• 1962. Review of F.F. Laidlaw & A. Solem ‘The land snail genus Amphidromus. A synoptic catalogue’. Journal of conchology 25: 87.
• 1962. Austroborus Parodiz a new name for Microborus Pilsbry. Journal of conchology 25: 133.
• 1962. Introduction to Molluscan taxonomy. The Conchologists' Newsletter 4: 20.
• 1962. Introduction to Molluscan taxonomy. The Conchologists' Newsletter 5: 24–25.

• & Crowley, T.E. 1963. Achatininae. In B. Verdcourt. The Miocene non-marine Mollusca of Rusinga Island, Lake Victoria and other localities in Kenya. Palaeontographica. Abteilung A 121 Abt. A:14-15. Reference page. 
• 1963. Pila letourneuxi (Bourguignat), its synonymy and distribution. Journal of conchology 25: 152–155, pl. 9.
• 1963. The Strophocheilidae: a neotropical family of terrestrial molluscs. The Conchologists' Newsletter 8: 41–43.

• & Beatty, D. 1964. A new species of freshwater gastropod Mollusca of the genus Saulea from the Miocene of Kenya. Breviora 212: 1–5, 1: 1–3.
• & Beatty, D. 1964. The Viviparidae. The Conchologists' Newsletter 11: 70–72.
• & Crowley, T.E. 1964. On Limicolariopsis d'hericourtiana (Bourguignat). Journal of conchology 25: 265–267 fig. Reference page. 
• & Crowley, T.E. 1964. Achatina (Lissachatina) tavaresiana Morelet; its synonymy and distribution. Revue de zoologie et de botanique africaines 69: 121–131, figs. Reference page. 
• & Crowley, T.E. 1964. Supplementary notes on the genus Limicolariopsis d'Ailly (Mollusca – Achatinidae). Revue de zoologie et de botanique africaines 69: 189–194, pl. 1. Reference page. 
• & Woodward, F.R. 1964. A monographic of the African bivalves of the genus Pleiodon Conrad (=Iridina authors) (Mollusca – Mutelidae). Annales du Musée Royal de l'Afrique Centrale: Serie in 8° Sciences Zoologiques 130: 1–33, pl. 1–4.
• 1964. The Pomacea flagellata complex in Central America. Journal of conchology 25: 224–231 pl. 13.
• 1964. A short glossary of molluscan terms. Conchological Society: Papers for Students 1964(4): 9 pp.
• T. E. Crowley & Woodward, F.R. 1964. A monographic review of the Mollusca of Lake Nyasa. Annales du Musée Royal de l'Afrique Centrale: Serie in 8° Sciences Zoologiques 131: 1–58, pl. 1–7, map.

• 1965. Land operculates of the New World. The Conchologists' Newsletter 15: 98–99.

• 1966. Distribution of the Acavidae. The Conchologists' Newsletter 12: 113–115. (1966).

• & Crowley, T.E. 1967. Further notes on the genus Limicolariopsis d'Ailly (Mollusca, Achatinidae). Revue de zoologie et de botanique africaines 75: 31–34. Reference page. 
• & Paul, C.R.C. 1967. Studies in the genus Pseudachatina Albers (Mollusca, Achatinidae). Annales du Musée Royal de l'Afrique Centrale: Serie in 8° Sciences Zoologiques 159: 1–79, pl. 1–5, text figures.
• & Preece, R.C. 1967. The land mollusca of the Bembridge limestone. Proceedings of the Isle of Wight Natural History and Archaeological Society 6: 101–111 (reprinted 1968).
• 1967. Some collecting experiences in British Guiana. The Conchologists' Newsletter 20: 139–140.

• & Woodward, F.R. 1968. A monograph of the African bivalves of the genera Brazzaea Bourguignat, Mweruella Haas, Prisodontopsis Tomlin and Pseudospatha Simpson. Revue de zoologie et de botanique africaines 77: 190–220, pl. 3: 4.
• 1968. Some collecting experiences in British Guiana ll. The Conchologists' Newsletter 24: 34–36.

• & Crowley, T.E. 1970. A monographic revision of the African land snails of the genus Limicolaria Schumacher (Mollusca – Achatinidae) Annales du Musée Royal de l'Afrique Centrale: Serie in 8° Sciences Zoologiques 77: 1–61, pl. 1–6. Reference page. 

• 1972. The Ampullariidae, an historical survey. Journal of conchology 27: 453–462.
• 1972. Achatina fulica Bowdich – a much travelled African Snail. The Conchologists' Newsletter 43: 280–281.

• & McMillan, N.F. 1974. A survey of the Mollusca of Lake Chad, Central Africa. Revue de zoologie africaine 88: 311–328, map. Reference page. 
• 1974. The land operculate genus Pollicaria Gould (Gastropoda), a systematic revision. Journal of conchology 28: 173–178 pl. 6.
• 1974. The genus Neptunea in Britain. The Conchologists' Newsletter 50: 390–391.

• 1976. Arthur Blok (1882–1974). Journal of conchology 29: 67–68.
• 1976. On the status of Murex osseus Reeve 1845 (Gastropoda Muricidae). Journal of conchology 29: 79–80.
• 1976. The Muricinae of the West African Marine Province. Newsletter of the British Shell Collectors' Club 1976(24): 17, 23 figs.
• 1976. Review of R.H. Fair ‘The Murex book’. The Conchologists' Newsletter 59: 458–459.
• 1976. The genus Neptunea in Britain. Ipswich Geological Group bulletin 1976(18): 9–11.

• & Crowley, T.E. 1977. Mollusca not Charopidae in La faune terrestre de l'île de Sainte-Hélène part 4. Annales du Musée Royal de l'Afrique Centrale: Serie in 8° Sciences Zoologiques 220: 534–575, pl. 9: 10. Reference page. 
• 1977. Review of G.E. Rodwin & A. D'Attiilio, ‘Murex shells of the World’. Journal of conchology 29: 224.
• 1977. On the occurrence of Colus ventricosus (Gray 1839), a prosobranch Mollusc. Bulletin of the Geological Society of Norfolk 29: 56–57.
• 1977. The genus Neptunea Roeding, 1798 in Western Europe (Prosobranchia – Buccinacea. La Conchiglia 9(101): 9–14, figs.

• & Crowley, T.E. 1978. A revision of the genus Revoilia Bourguignat 1881 (Prosobranchia: Pomatiidae). Journal of conchology 29: 351–364 pl8-9. Reference page. 
• & Edwards, N. 1978. On the status of Potomya J. de C. Sowerby 1835 (Bivalvia: Myoida). Journal of conchology 29: 301–303.
• 1978. How rare is Murex barclayi Reeve? Newsletter of the British Shell Collectors' Club 1978(32): 5.
• 1978. The genus Colus Roeding, 1798 in western Europe (Prosobranchia – Buccinoidea. La Conchiglia 10(114-115): 3–7, fig1-13.

• 1979. Siphonorbis ebur (Mörch) (Buccinidae) and its occurrence in British waters. Newsletter of the British Shell Collectors' Club 1979(37): 9–10, figs.
• 1979. Siphonorbis Mörch, 1869: a deep water genus from the North Atlantic (Buccinidae – Prosobranchia). La Conchiglia 11(118-119): 6–7, figs.
• 1979. The genus Buccinum Linné, 1758 in western Europe (Prosobranchia – Buccinnoidea) Part 1: from the British seas to the Mediterranean. La Conchiglia 11(126-127): 15–18, figs.
• 1979. Review of Wagner & Abbott's ‘Standard Catalogue of shells supplement 1’. The Conchologists' Newsletter 68: 130–131.

• 1980. More about Achatina. Hawaiian Shell News 28(10): 7.
• 1980. The Buccinids from European seas. La Conchiglia 12(138-139): 16–18, figs.

• & Crowley, T.E. 1981. Achatina (Lissachatina) mulanjensis a new species from Malawi (Mollusca, Gastropoda, Pulmonata). Revue de zoologie africaine 95: 954–958, 6. Reference page. 
• 1981. Meet the 'Strophocheilidae. Hawaiian Shell News 29(8): 1, 6, figs.
• 1981. The genus Vitularia Swainson, 1840 (Prosobranchia – Muricidae). La Conchiglia 13(142-143): 8–9, figs.
• 1981. Troschelia berniciensis (King, 1876) (Fasciolariidae – Fusininae). La Conchiglia 13(148-149): 3–4, figs.
• 1981. The genus Purpurellus Jousseaume, 1880 (Gastropoda: Muricidae). La Conchiglia 13(148-149): 4–5, figs.
• 1981. Review of R. Houart ‘Révision des Trophoninae d'Europe’. The Conchologists' Newsletter 77: 319.

• Houart, R. & 1982. On the description of a neotype for Chicoreus (Chicoreus) torrefactus (Sowerby Jr., 1841) and description of a new species: Chicoreus (Chicoreus) kilburni sp. nov. (Gastropoda: Muricidae: Muricinae). Informations de la Société belge de malacologie 10: 51–56. Reference page. 
• 1982. The genus Buccinum Linn. 1758 in western Europe (Prosobranchia: Buccinidae). Part 2: European arctic species. La Conchiglia 14(158-159): 14–18 figs.
• 1982. Colus solidus Sowerby, 1880 (Prosobranchia: Buccinidae). A little known N. Atlantic species. La Conchiglia 14(160-161): 3 figs.
• 1982. On the occurrence of Colus ventricosus Gray 1839 a prosobranch mollusc. The Conchologists' Newsletter 80: 369.

• & Houart, R. 1983. The Chicoreus (Chicoreus) torrefactus complex Part 2. La Conchiglia 15(170-171): 3–6, figs. Reference page. 
• & Oliver, A.P.H. 1983. Is Bursa elegans Sowerby 1841 a good species? Newsletter of the British Shell Collectors' Club 14(2): 30, fig.
• 1983. Marine shells from an African lake. Hawaiian Shell News 31(1): 1, 4, figs.
• 1983. Is this the most ponderous land snail? Hawaiian Shell News 31(10): 3, fig.
• 1983. Marine species from a freshwater lake? Newsletter of the British Shell Collectors' Club 14(2): 17–18 figs.
• 1983. The Chicoreus (Chicoreus) torrefactus complex Part 1. La Conchiglia 15(168-169): 16–19, figs.

• & Kermarrec-Labisse, A. 1984. An unusual form of southern Iceland Neptunea despecta fornicata (Fabricius, 1780). La Conchiglia 16(180-181): 19, figs.
• 1984. Placostylus senilis (Gassies, 1869). Is this the worlds most ponderous landshell? Newsletter of the British Shell Collectors' Club 14 (4) 1984(56): 59.
• 1984. The genus Pyrolofusus Mörch, 1869 (Prosobranchia: Buccinidae). La Conchiglia 16(178-179): 6–7, figs.

• 1985. The genus Japelion Dall, 1916 (Prosobranchia: Buccinidae). La Conchiglia 17(194-195): 24–27, figs.

• & Nelson, C.M. 1986. Linnaeus' Neptunea, (Mollusca: Gastropoda) Zoological Journal of the Linnean Society 88: 291–306, 6 figs.

• 1987. The genus Clinopegma Grant & Gala, 1931. La Conchiglia 19(222-223): 3, figs.
• 1987. The saga of the turban star shell Megastraea turbanica (Dall). Pallidula 18(2): 19–21.
• 1987. Corrections to the first printing of Abbott & Dance, ‘Compendium of Seashells’ 1982 Family Bursidae. Pallidula 18(3): 37.

• 1988. A new look at Echinophoria Sacco, 1890. La Conchiglia 1988(236-237): 19 figs.

• 1989. Rangia cuneata, the clam they built a town on. Pallidula 19(4): 62–63.
• 1989. Cassis nana and thompsoni: range extensions. Pallidula 19(4): 73.

• & Cox, G.H.J. 1990. Cassis bituberculosa v. Mts. 1901 a species of Echinophoria Sacco, 1890. La Conchiglia 22(250-252): 6–7, fig.
• 1990. Ceratostoma coreanicum (A. Adams, 1854) a synonym of C. burnetti Adams & Reeve 1849. La Conchiglia 22(256): 22, figs.
• 1990. The saga of the ‘blubber-lipped bulimus’. Pallidula 20(2): 21.

• 1991. Pleurotomaria: 225 million years of perseverance? Pallidula 21(2): 7–10.

• 1992. The genus Megastraea McLean, 1970 (Archaegastropoda: Turbinidae). La Conchiglia 23(263): 4, 4 figs.
• 1992. Gordon Osbom F.L.S., 1913-1991. Pallidula 22(1): 5.
• 1992. A brief look at ammonites. Pallidula 22(1): 16–19.

• 1993. Review of I. Bogdanov and B. Sirenko ‘Seashells of Russia in colour’. Pallidula 23(2): 26–27.

• 1994. The giant creeper Campanile symbolicum Iredale a relict from Australian waters. Pallidula 24(1): 18–20.

• 1995. A living fossil from the southwest Pacific Gourmya gourmyi (Crosse). Pallidula 25(1): 18–20.

• 1996. Neotrigonia, a sole survivor! Pallidula 26(1): 22–23.
• 1996. The repetitive shell name once again. Pallidula 26(1): 25.

• 1997. The killer sea snail Conus geographus Linnaeus, 1758. Pallidula 27(1): 17–18.

• 1998. Our common whelk Buccinum undatum Linné, 1758. Pallidula 28(1): 6–7.

• & Pain, C. 2000. Terence Eldon Crowley 1915–1999: obituary, bibliography and new molluscan names. Journal of conchology 37: 93–97. BHL Reference page. 

• Achatina bandeirana arenaria Crowley & Pain 1961.
• Achatina bandeirana mayumbensis Crowley & Pain 1961.
• Achatina (Lissachatina) calcicola Crowley & Pain 1961.
• Achatina (Pintoa) lomaniensis Crowley & Pain 1961.
• Achatina (Lissachatina) mulanjensis Crowley & Pain 1981.
• Achatina schweinfurthi simularis Crowley & Pain 1961.
• Archachatina (Tholachatina) altitudinaria Crowley & Pain 1961.
• Archachatina (Tholachatina) bequaerti Crowley & Pain 1961.
• Archachatina (Tholachatina) insularis Crowley & Pain 1961.
• Burtoa nilotica benoiti Crowley & Pain 1959.
• Burtoa nilotica congoensis Crowley & Pain 1959.
• Burtoa nilotica verdcourti Crowley & Pain 1959.
• Chicoreus (Chicoreus) kilburni Houart & Pain 1982.
• Grandidieria bloomeri Woodward in Crowley, Pain & Woodward 1964.
• Lanistes (Meladomus) connollyi Pain 1954.
• Lanistes kobelti Pain 1954.
• Lanistes (Meladomus) ovum mweruensis Pain 1954.
• Limicolaria leakeyi Crowley & Pain 1963.
• Limicolariopsis cylindrica Crowley & Pain 1961.
• Limicolariopsis elgonensis Crowley & Pain 1961.
• Limicolariopsis ellisi Crowley & Pain 1964.
• Limicolariopsis laevis Crowley & Pain 1961.
• Limicolariopsis verdcourti Crowley & Pain 1961.
• Neothauma ecclesi Crowley, Pain & Woodward, 1964.
• Nyassunio ecclesi Woodward in Crowley, Pain & Woodward 1964.
• Pila (Pomacea) aldersoniPain 1946.
• Pila ovata dartevellei Pain 1961.
• Pila (Pomacea) hollingsworthi Pain 1946.
• Placostylus fibratus kumacensis Pain
• Placostylus fibratus mosesi Pain 1955.
• Placostylus fibratus powelli Pain 1955.
• Placostylus scarabus verdis Pain 1955.
• Placostylus strattoni Pain 1955.
• Pleiodon waterstoni Pain & Woodward 1964.
• Pomacea camena Pain 1949.
• Pomacea cumingi urabaensis Pain 1956.
• Pomacea falconensis Pain & Arias 1958.
• Pomacea (Limnopomus) meridaensis Pain 1950.
• Pomacea nais Pain 1949.
• Pomacea vickeryi Pain 1949.
• Pomacea zischkai Blume & Pain 1952
• Pseudachatina connectens d’Aily forma colorata Pain & Paul 1967.
• Pseudachatina connectens d’Ailly forma plicata Pain & Paul 1967.
• Pseudachatina granulata Pain & Paul 1967.
• Pseudachatina pulchra Pain & Paul 1967.
• Pseudachatina sodeni Kobelt forma elegantula Pain & Paul 1967.
• Pseudachatina striata Pain & Paul 1967.
• Pseudachatina wrighti (G.B. Sow. sen.) forma minor Pain & Paul 1967
• Saulea lithoides Pain & Beatty 1964.

• Conchological Society of Great Britain & Ireland: Verdcourt, A.H. Wood, and B. Rowson - Thomas Pain, 1915–2003