Keith Richard Willmott

Keith Richard Willmott, English entomologist. Author abbreviation: Willmott

• McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA

(List may be incomplete)

• 129 taxon names authored by Keith Richard Willmott

(List may be incomplete)

• 1 eponym of Keith Richard Willmott

(List may be incomplete)

• & Hall, J.P.W. 1994. Four new species of Riodinids from western Ecuador (Lepidoptera: Riodinidae). Tropical lepidoptera 5(2): 87-91. Full PDF. Reference page. 

• Hall, J.P.W. & , 1995. Two new species of Adelpha from eastern Ecuador (Lepidoptera: Nymphalidae). Tropical Lepidoptera 6 (2): 106–109. Full article: [1]. Reference page. 
• Hall, J.P.W. & 1995a: Notes on the genus Argyrogrammana, with descriptions of five new species (Lepidoptera: Riodinidae). Tropical lepidoptera 6(2): 136–143. Full article (pdf). Reference page. 
• Hall, J.P.W. & , 1995b. Five new species and a new genus of riodinid from the cloud forests of eastern Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera Research 6(2): 131–135. Full article: PDF. Reference page. 
• Hall, J.P.W. & 1995c. Two new species of Mesene from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera 6(2): 110-112. PDF. Reference page. .
• Hall, J.P.W. & 1995d. Two new species of satyrines from Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera 6(2): 103-105. PDF. Reference page. 

• Hall, J.P.W. & 1996. Systematics of the riodinid tribe Symmachiini, with the description of a new genus and five new species from Ecuador, Venezuela and Brazil (Lepidoptera: Riodinidae). Lambillionea 96(4): 637–660. PDFa PDFb Reference page. 
• Hall, J.P.W. & 1996. The genus Theope: four new species and a new subspecies (Lepidoptera: Riodinidae). Tropical lepidoptera 7(1): 63–67. Abstract (pdf). Reference page. 
• Hall, J.P.W. & , 1996. Notes on the genus Argyrogrammana. Part 2, with one new species (Lepidoptera: Riodinidae). Tropical Lepidoptera 7 (1): 71–80. Abstract: [2]. Reference page. 
• Torres-Núñez, R., Hall, J.P.W., & Johnson, K. 1996. A new genus of Elfin butterflies from the northern high Andes (Lepidoptera: Lycaenidae). Tropical lepidoptera 7(1):81-86.Reference page. 

• Hall, J.P.W.; 1998: Four new Riodinid species from eastern Ecuador (Lepidoptera: Riodinidae). Lambillionea 98(3): 325–334. Full article Reference page. 
• Hall, J.P.W. & 1998. Nine new species and one new subspecies of Euselasia from Ecuador (Lepidoptera: Riodinidae). Tropical lepidoptera 9 supplement 1: 27–35. full article (pdf). Reference page. 
• Hall, J.P.W. & 1998b. Three new species of Riodinini from the cloud forests of Ecuador. (Lepidoptera: Riodinidae). Tropical lepidoptera 9 supplement 1: 22–26. Full article (pdf). Reference page. 

• Pyrcz, T.W., & Hall, J.P.W., 1999: Contribution to the knowledge of Ecuadorian Pronophilini, Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae). Genus 10(3): 497-522. full article (pdf)Reference page. 

• , Hall, J.P.W. & Lamas, G. 2001. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes. Systematic Entomology 26: 369–399. PDF Reference page. 

• & Mallet, J. 2004. Correlations between adult mimicry and larval hostplants in ithomiine butterflies. Proceedings of the Royal Society of London B (Biology Letters Suppl.) 271: S266-S269. Reference page. 

• Hall, J.P.W.; , Busby, R.C. 2005: Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru. Zootaxa 797: 1–20. Abstract & excerpt Reference page. 
• Hall, J. & , 2005. A New Species of Paiwarria (Lepidoptera: Lycaenidae: Eumaeini) from Western Ecuador. Proceedings of the Entomological Society of Washington 107 (4): 960–967. BHL. Reference page. 

• & Freitas, A.V.L. 2006. Higher-level phylogeny of the Ithomiinae (Lepidoptera: Nymphalidae): classification, patterns of larval hostplant colonisation and diversification. Cladistics 22: 297-368. Reference page. 
• & Lamas, G. 2006. A phylogenetic reassessment of Hyalenna Forbes and Dircenna Doubleday, with a revision of Hyalenna (Lepidoptera: Nymphalidae: Ithomiinae). Systematic Entomology 31(3): 419-468. Reference page. 

• Hall & Willmott, [2007]. Four new species of Symmachiini from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera 16 (1-2): 1–5. PDF. Reference page. 
• & Lamas, G. 2007. A revision of Pachacutia, a new genus of rare Andean ithomiine butterflies (Nymphalidae: Ithomiinae), with the description of two new species. Annals of the Entomological Society of America 100: 449-469. DOI: 10.1603/0013-8746(2007)100[449:AROPAN]2.0.CO;2. . Reference page. 

• & Vitale, F. 2008. Taxonomic notes on Napeogenes (Lepidoptera: Ithomiinae) from Ecuador and Colombia, with the description of ten new subspecies. Genus 19(3): 585-609. Reference page. 
• & Lamas, G. 2008. A revision of the genus Megoleria (Lepidoptera: Nymphalidae: Ithomiini, (Ithomiina), Tropical Lepidoptera Research 18(1): 36-59 (PDF). Reference page. 

• Hall, J.P.W. & , 2009. Two new species of Euselasia (Riodinidae: Euselasiinae) from Western Ecuador. Tropical Lepidoptera Research 19(1): 52–55. PDF Reference page. 

• , Elias, M. & Sourakov, A. 2011. Two Possible Caterpillar Mimicry Complexes in Neotropical Danaine Butterflies (Lepidoptera: Nymphalidae). Annals of the Entomological Society of America 104(6): 1108-1118. DOI: 10.1603/AN10086. . Reference page. .

• Nakahara, S., Hall, J.P.W., Lamas, G. & Willmott, K.R. 2015. Seven new species and one new subspecies of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the tropical Andes. Tropical Lepidoptera Research 25(2): 63–79. Reference page. 
• 2015. A new subspecies of Cithaerias pyropina (Salvin & Godman, 1868) from the Cordillera de Cutucú in southeastern Ecuador (Nymphalidae: Satyrinae). Tropical lepidoptera Research 25(1): 1-4. full article (pdf). Reference page. 

• Freitas, A.V.L., Barbosa, E.P., , Wahlberg, N. & Lamas, G. 2016. ‘Species’ from two different butterfly genera combined into one: description of a new genus of Euptychiina (Nymphalidae: Satyrinae) with unusually variable wing pattern. Revista Brasileire de Entomologia 60(3): 157–165. full article (PDF). Reference page. 
• Freitas, A.V.L., Magaldi, L.M. & 2016. A new subspecies of Oleria gunilla (Nymphalidae: Danainae) from North Mato Grosso, Brazil. Journal of the Lepidopterists’ Society 70(4): 295–301 DOI: 10.18473/lepi.70i4.a8. Reference page. 
• Nakahara, S., Tan, D., Lamas, G., Parus, A. & 2016. A distinctive new species of Hermeuptychia Forster, 1964 from the eastern tropical Andes (Lepidoptera: Nymphalidae: Satyrinae) Tropical Lepidoptera Research 26(1): 6-12. full article (PDF). Reference page. 
• Nakahara, S., Barbosa, E.P., Marin, M.A., Freitas, A.V.L., Pomerantz, T. & 2016a. Graphita gen. nov., a New Genus for Neonympha griphe C. Felder & R. Felder, 1867. (Lepidoptera, Nymphalidae, Satyrinae). Neotropical Entomology 45: 675–691. DOI: 10.1007/s13744-016-0414-x. . Reference page. 
• Nakahara, S., Vega, G. & Willmott, K.R. 2016. Description of a new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the western Andes. Zootaxa 4184(2): 358–366. DOI: 10.11646/zootaxa.4184.2.7. Reference page. 
• , Mielke, O.H.H. & Freitas, A.V.L. 2016. A new subspecies and validation of the enigmatic species Adelpha herbita Weymer endemic to the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 26(1): 32-37. full article (PDF). Reference page. 

• , Lamas, G. & Hall, J.P.W. 2017. Notes on the taxonomy of Actinote intensa Jordan (Lepidoptera: Nymphalidae: Heliconiinae) and the description of a new sibling species from eastern Ecuador. Tropical Lepidoptera Research 27(1): 6-15. full article (PDF). Reference page. 
• , Robinson Willmott, J.C., Elias, M. & Jiggins, C.D. 2017a. Maintaining mimicry diversity: optimal warning color patterns differ among microhabitats in Amazonian clearwing butterflies. Proceedings of the Royal Society B 284(1855)(20170744): 1-9. DOI: 10.1098/rspb.2017.0744. . Reference page. 
• , Simon, M., Ortiz-Acevedo, E. & Hall, J.P.W. 2017. First record of the enigmatic tribe Anaeomorphini (Lepidoptera, Nymphalidae, Charaxinae) outside of the Amazon basin: a new species of Anaeomorpha Rothschild, 1894, from the Chocó region of western Ecuador. Insecta Mundi (0548): 1–10. Full article (PDF). Reference page. 

• Barbosa, E.P., Siewert, R.R., Mielke, O.H.H., Lamas, G., Willmott, K.R. & Freitas, A.V.L.. 2018. Redescription of Yphthimoides patricia (Hayward, 1957), with taxonomic notes on the names Euptychia saltuensis Hayward, 1962 and Yphthimoides manasses (C. Felder & R. Felder, 1867) (Nymphalidae: Satyrinae). Zootaxa 4422(4): 537–557. DOI: 10.11646/zootaxa.4422.4.5 Reference page. 
• Benmesbah, M., Zacca, T., Casagrande, M.M., Mielke, O.H.H., Lamas, G. & 2018. Taxonomic notes on Papilio ocypete Fabricius, 1776 and Papilio helle Cramer, 1779 with description of two new similar species from South America (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 4425(1): 115–145. DOI: 10.11646/zootaxa.4425.1.7. Reference page. 
• Espeland, M., Breinholt, J.W., , Warren, A.D., Vila, R., Toussaint, E.F.A., Maunsell, S.C., Aduse-Poku, K., Talavera, G., Eastwood, R.G., Jarzyna, M.A., Guralnick, R., Lohman, D.J., Pierce, N.E. & Kawahara, A.Y. 2018. A Comprehensive and Dated Phylogenomic Analysis of Butterflies. Current Biology 28(5): 770–778. DOI: 10.1016/j.cub.2018.01.061 Reference page. 
• Nakahara, S., Zacca, T., Huertas, B., Neild, A.F.E., Hall, J.P.W., Lamas, G., Holian, L.A., Espeland, M. & 2018: Remarkable sexual dimorphism, rarity and cryptic species: a revision of the ‘aegrota species group’ of the Neotropical butterfly genus Caeruleuptychia Forster, 1964 with the description of three new species (Lepidoptera, Nymphalidae, Satyrinae). Insect Systematics & Evolution 49(2): 130–182. DOI: 10.1163/1876312X-00002167 . Reference page. 
• Nakahara, S., , Mielke, O.H.H., Schwartz, J., Zacca, T., Espeland, M. & Lamas, G. 2018a: Seven new taxa from the butterfly subtribe Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) with revisional notes on Harjesia Forster, 1964 and Pseudeuptychia Forster, 1964. Insecta Mundi 0639: 1–38. Reference page. 
• , Lamas, G., Radford, J.T., Marín, M.A., Nakahara, S., Espeland, M., Xiao, L. & Hall, J.P.W. 2018. A distinctive new species of cloud forest Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from Ecuador and Peru. Tropical Lepidoptera Research 28(1): 39-45. full article (PDF). Reference page. 

• Freitas, Woodbury, M.S., Barbosa & 2019. A new subspecies of Adelpha messana (C. Felder & R. Felder) from the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 29(2): 62-66. Online. Reference page. .
• Nakahara, S., Lamas, G., Tyler, S., Marín, M.A., Huertas, B., , Mielke, O.H.H. & Espeland, M. 2019: A revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilio arnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae). ZooKeys 821: 85–152. Reference page. 
• , Marín, M.A., Nakahara, S., Pomerantz, T., Lamas, G., Huertas, B., Espeland, M., Xiao, L., Hall, J.P.W., Willmott, J.I.R. & Freitas, A.V.L. 2019. A revision of the new Andean butterfly genus Optimandes Marín, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research 29(1): 29-44. Online. Reference page. .

• Nakahara, S., Kleckner, K., Lamas, G., Huertas, B. & 2020. A contribution towards the systematics of Magneuptychia Forster, 1964: Caeruleuptychia francisca (Butler, 1870), n. comb. (Lepidoptera: Nymphalidae: Satyrinae). Tijdschrift voor Entomologie 163(1): 51-61. Reference page. 
• , Lamas, G., Hall, J.P.W., Mota, L.L. & Kell, T. 2020. The common, the rare, and the lost: Descriptions of twelve new species and three new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research 30(1): 1-49. Reference page. 
• Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Espeland, M., Freitas, A.V.L., , Nakahara, S. & Lamas, G. 2020. Revalidation of Vareuptychia Forster, 1964, description of Vanima gen. nov., and notes on Euptychia cleophes Godman & Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 4858(1): 1–34. DOI: 10.11646/zootaxa.4858.1.1. . Reference page. 
• Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Barbosa, E.P., Freitas, A.V.L. & 2020a. Description of Emeryus Zacca, Mielke & Casagrande gen. nov. (Lepidoptera: Nymphalidae) to accommodate three species formerly placed in Paryphthimoides Forster, 1964. Austral Entomology 59: 505–523. DOI: 10.1111/aen.12479. Reference page. 

• Mahecha-Jiménez, O., Florczyk, K., , Cerdeña, J.A., Zubek, Boyer, P., Farfán, J., Lachowska-Cierlik, D. & Pyrcz, T.W. 2021. Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru: description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina). Zootaxa 5072(3): 201–237. DOI: 10.11646/zootaxa.5072.3.1. . Reference page. 
• Nakahara, S., Matos-Maraví, P., Schwartz, J. & 2021. Assessing a generic synapomorphy of Pseudodebis Forster, 1964 (Lepidoptera : Nymphalidae : Satyrinae) and a recent speciation with a shift in elevation between two new species in the western Andes. Invertebrate Systematics 35: 158–180. DOI: 10.1071/IS20024. . Reference page. 
• Nakahara, S., Matos-Maraví, P., , Nakamura, I. & MacDonald, J.R. 2021. Description of a new species of Pseudodebis Forster, 1964 from Central America (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research 31(1): 35-41. Reference page. 
• Nakahara, S., , MacDonald, J.R. & Thurman, A. 2021b. Description of an enigmatic new species of Chloreuptychia Forster, 1964 from Panama (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research 31(3): 145-150. Reference page. 
• Neild, A.F.E., Losada, M.E. & 2021. A distinctive new subspecies of Moschoneura pinthous (Linnaeus, 1758) (Lepidoptera: Pieridae: Dismorphiinae) from the Orinoco Delta, Venezuela, with comments on the species-level taxonomy. Tropical Lepidoptera Research 31(1): 60-67. Reference page. 
• Padrón, P.S., Pyrcz, T.W. & 2021. A Contribution Towards Resolving the Systematics of the High-Altitude Tropical Andean Satyrine Genus Altopedaliodes Forster, 1964 (Lepidoptera, Nymphalinae: Satyrinae). Neotropical Entomology 50: 767–803. DOI: 10.1007/s13744-021-00897-9. . Reference page. 
• Rosa, A.H.B., Huertas, B., , Barbosa, E.P., Machado, P.A., Mielke, O.H.H., Canaan, C.H.P. & Freitas, A.V.L. 2021. Fifty years without a name: a new species of Splendeuptychia Forster (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5061(1): 095–114. DOI: 10.11646/zootaxa.5061.1.4. . Reference page. 
• Willmott, K.R., Lamas, G., Hall, J.P.W., Vitale, F., Boyer, P., Petit, J.C., Radford, J.T. & Elias, M. 2021. A new species and thirty-eight new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research 31 (Supplement 3): 1-80. DOI: 10.5281/zenodo.5526837. Reference page. 
• Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Barbosa, E.P., Freitas, A.V.L.. Lamas, G., Espeland, M., Brévignon, C., Nakahara, S., Checa, M.F. & 2021. Systematics of the Neotropical butterfly genus Paryphthimoides Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae), with descriptions of seven new taxa. Insect Systematics & Evolution 52(1): 42–96. DOI: 10.1163/1876312X-00001027 . Reference page. 
• Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Freitas, A.V.L.. Marín, M.A., Espeland, M. & 2021a. A new euptychiine butterfly species from south Brazil and taxonomic rearrangements for Taydebis Freitas, 2013 and Hermeuptychia Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5023(4): 555–570. DOI: 10.11646/zootaxa.5023.4.5. . Reference page. 

• Mota, L.L., Rosa, A.H.B., Vasconcellos, L.R., & Freitas, A.V.L. 2022. A new subspecies of Mechanitis lysimnia from southern Amazonia (Nymphalidae: Danainae: Ithomiini). Tropical Lepidoptera Research 32(1): 47-51. Reference page. 

• Hall, J.P.W. & 2023. A review of the Argyrogrammana fauna of the Transandean region (Lepidoptera: Riodinidae: Symmachiini). Tropical Lepidoptera Research 33(supplement 1): 28-48. Reference page. 
• Hall, J.P.W., & Ahrenholz, D.H. 2023. A review of the Argyrogrammana fauna of the eastern Andes (Lepidoptera: Riodinidae: Symmachiini) Tropical Lepidoptera Research 33(supplement 1): 49-60. Reference page. 
• Hall, J.P.W., & Ahrenholz, D.H. 2023a. A review of the Argyrogrammana amalfreda group fauna of the western Amazon (Lepidoptera: Riodinidae: Symmachiini). Tropical Lepidoptera Research 33(supplement 1): 61-85. Reference page. 
• Nakahara, S., Kleckner, K., Barbosa, E.P., Lourenço, G.M., Casagrande, M.M., Willmott, K. & Freitas, A.V.L. 2023. Reassessment of the type locality of Euptychia stigmatica Godman, 1905, with the description of two new sibling species from Amazonia (Lepidoptera, Nymphalidae, Satyrinae, Satyrini). Zookeys 1167ː 57–88. DOI: 10.3897/zookeys.1167.102979 Reference page. 

• Pyrcz, T.W., Boyer, P., Petit, J.C., Garlacz, R., Zajac-Garlacz, K.S., Espeland, M. & 2024. Bedazzled: a new, striking species of Corades from the outskirts of Quito questions our knowledge of Andean cloud forest butterflies (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa 5453(2): 255–262. DOI: 10.11646/zootaxa.5453.2.6. Reference page. 
• , Lamas, G., Hall, J.P.W., Boyer, P., Pyrcz, T.W. & Florczyk, K. 2024. A new species and eight new subspecies of high elevation Actinote from Colombia, Ecuador and Peru (Nymphalidae, Heliconiinae, Acraeini). Tropical Lepidoptera Research 34(1): 1-20. Reference page. 

• List of publications. 2016. Systematics of Neotropical Butterflies. McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History.