You can use traceback() to locate where the last error occurred. Usually it will point you to a call you make in your function. Then I typically put browser() at that point, run the function again and see what is going wrong.
For example, here are two functions:
f2 <- function(x)
{
if (x==1) "foo"
}
f <- function(x)
{
f2(x)
}
Note that f2() assumes an argument of length 1. We can misuse f:
> f(NULL)
Error in if (x == 1) "foo" : argument is of length zero
Now we can use traceback() to locate what went wrong:
> traceback()
2: f2(x) at #3
1: f(NULL)
The number means how deep we are in the nested functions. So we see that f calls f2 and that gives an error at line 3. Pretty clear. We could reassign f with browser placed just before the f2 call now to check it's input. browser() simply allows you to stop executing a function and look around in its environment. Similar to debug and debugonce except that you don't have to execute every line up until the point you know something goes wrong.