I've looked for an answer but I still have not found a solution, I'm still new to R. My data frame displays measurement of one ecological trait (relative soil coverage) for ~70 plant species in different conditions: different years, different chemical treatment and presence/absence of greenhouse.
I need to summarize that data into a new dataframe that displays the mean and sd of the trait for each species and for each combination of factors (conditions). I know aggregate or lapply could help but i struggle to combine both grouping by 3 different factors and the multiples species, which implies the need of an "automated" code. 
I'm sorry if I missed a post answering my question
Thank you for your patience and help
Edit: here is a reproducible example, hope i do that correctly:
mydata<-structure(list(Year = c(2010L, 2010L, 2010L, 2010L, 2010L, 2010L, 
2010L, 2010L, 2011L, 2011L, 2011L, 2011L, 2011L, 2011L, 2011L, 
2011L), Replicate = c(1L, 1L, 1L, 1L, 2L, 2L, 2L, 2L, 1L, 1L, 
1L, 1L, 2L, 2L, 2L, 2L), Treatment = structure(c(1L, 2L, 1L, 
2L, 1L, 2L, 1L, 2L, 1L, 2L, 1L, 2L, 1L, 2L, 1L, 2L), .Label = c("A", 
"B"), class = "factor"), Greenhouse = structure(c(2L, 2L, 1L, 
1L, 2L, 2L, 1L, 1L, 2L, 2L, 1L, 1L, 2L, 2L, 1L, 1L), .Label = c("No", 
"Yes"), class = "factor"), Sp_1 = c(4L, 0L, 2L, 5L, 4L, 0L, 2L, 
5L, 0L, 0L, 4L, 6L, 4L, 0L, 2L, 5L), Sp_2 = c(7L, 0L, 1L, 1L, 
7L, 0L, 1L, 1L, 7L, 0L, 1L, 1L, 6L, 0L, 1L, 1L), Sp_3 = c(8L, 
2L, 2L, 1L, 8L, 2L, 2L, 1L, 10L, 2L, 1L, 1L, 4L, 2L, 2L, 1L)), class = "data.frame", row.names = c(NA, 
-16L))
I only put 3 species in that example but as i've said i have over 70 species so i need something that could select all the species columns ( mydata[,5:75] ? something along those lines) more than c("sp_1","sp_2",..., "sp_70"). 
I'd like the output to look like this:
Year   Treatment   Greenhouse   Sp_1_mean   Sp_1_sd   Sp_2_mean   Sp_2_sd 
2010   A           Yes          x           x         x           x
2010   A           No           x           x         x           x
2010   B           Yes          x           x         x           x
2010   B           No           x           x         x           x
2011   A           Yes          x           x         x           x
2011   A           No           x           x         x           x
2011   B           Yes          x           x         x           x
2011   B           No           x           x         x           x
Here is a dput() showing what the desired output should look like
    desired_output<-structure(list(Year = c(2010L, 2010L, 2010L, 2010L, 2011L, 2011L, 
2011L, 2011L), Treatment = structure(c(1L, 1L, 2L, 2L, 1L, 1L, 
2L, 2L), .Label = c("A", "B"), class = "factor"), Greenhouse = structure(c(2L, 
1L, 2L, 1L, 2L, 1L, 2L, 1L), .Label = c("No", "Yes"), class = "factor"), 
    Sp_1_mean = structure(c(1L, 1L, 1L, 1L, 1L, 1L, 1L, 1L), .Label = "x", class = "factor"), 
    Sp_1_sd = structure(c(1L, 1L, 1L, 1L, 1L, 1L, 1L, 1L), .Label = "x", class = "factor"), 
    Sp_2_mean = structure(c(1L, 1L, 1L, 1L, 1L, 1L, 1L, 1L), .Label = "x", class = "factor"), 
    Sp_2_sd = structure(c(1L, 1L, 1L, 1L, 1L, 1L, 1L, 1L), .Label = "x", class = "factor"), 
    Sp_3_mean = structure(c(1L, 1L, 1L, 1L, 1L, 1L, 1L, 1L), .Label = "x", class = "factor"), 
    Sp_3_sd = structure(c(1L, 1L, 1L, 1L, 1L, 1L, 1L, 1L), .Label = "x", class = "factor")), class = "data.frame", row.names = c(NA, 
-8L))
I hope that's clearer! Thanks
 
    