Osvaldo Villarreal-Manzanilla

Osvaldo Villarreal also published as Osvaldo Villarreal-Manzanilla/ Osvaldo Villarreal M., Venezuelan arachnologist & herpetologist.

Departamento de Invertebrados, Museu Nacional/UFRJ, Quinta da Boa Vista, São Cristóvão, 20.940-040 Rio de Janeiro, RJ, Brazil.

(List may be incomplete)

• 13 taxon names authored by Osvaldo Villarreal-Manzanilla

(List may be incomplete)

Link to: Overview of Arachnology

• Villarreal Manzanilla, O. 2004. A new species of the genus Stygnidius (Opiliones, Stygnidae) from Mount Marahuaca in Amazonas state, Venezuela. Revista Ibérica de Aracnología 9: 15–18. PDF. Reference page. 

• & Pinto-da-Rocha, R. 2006. Five new species of Protimesius from Brazil (Opiliones: Stygnidae). Zootaxa 1325: 219–233. Abstract & excerpt. Reference page. 

• Kury, A.B., Villarreal M, O. & Sampaio [Costa], C. 2007. Redescription of the type species of Cynorta (Arachnida, Opiliones, Cosmetidae). Journal of Arachnology 35(2): 325-333. DOI: 10.1636/H06-35.1. PDF . Reference page. 

• Pinto-da-Rocha, R. & Villarreal M, O. 2009. Cladistic analysis of the Stygninae and description of a new species of Protimesius Roewer, 1913 (Opiliones: Stygnidae). Zootaxa 2176: 48–56. Abstract & excerpt . Reference page. 
• Villarreal M, O. & Kury, A.B. 2009. A new generic homonym in the Agoristenidae (Arachnida: Opiliones). Zootaxa 2045: 65–68. Abstract & excerpt . Reference page. 

• Villareal M., O. & Kury, A.B. 2012. Licornus Roewer, 1932: newly transferred to Ampycinae and first record of the family Gonyleptidae (Opiliones: Laniatores) from Venezuela. Zootaxa 3544: 71–78. Preview Reference page. 

• Hara, M.R., Pinto-da-Rocha, R. & Villarreal M., O. 2014. Revision of the cranaid genera Phalangodus, Iquitosa and Aguaytiella (Opiliones: Laniatores: Gonyleptoidea). Zootaxa 3814(4): 567–580. DOI: 10.11646/zootaxa.3814.4.8 Reference page. 

• Villarreal M, O., Kury, A.B. & Pinto-da-Rocha, R. 2015. The poorly known genus Ventrifurca Roewer, 1913 revisited (Opiliones: Cranaidae). Zoological Studies 54(45): 1–18. Full article (PDF). Reference page. 

• Giupponi, A.P. de L., Miranda, G.S. de, Villarreal, O.M. 2016. Rowlandius dumitrescoae species group: new diagnosis, key and description of new cave-dwelling species from Brazil (Schizomida, Hubbardiidae). ZooKeys 632: 13–34. DOI: 10.3897/zookeys.632.9337. Reference page. 
• Indicatti, R.P. & Villarreal, O.M. 2016: Pseudonemesia tabiskey, a new species of Pseudonemesia Caporiacco 1955 and new ultramorphological data for the Microstigmatinae (Araneae: Microstigmatidae). Journal of Natural History 50(33/34): 2153-2167. DOI: 10.1080/00222933.2016.1196297. Reference page. 

• García, A.F. & Villarreal M, O. 2020. Description of a new species of Leptostygnus Mello-Leitão, 1940 and notes on the male genitalia in the subfamily Leiosteninae (Opiliones: Agoristenidae). Studies on Neotropical Fauna and Environment 55(3): 165–179. DOI: 10.1080/01650521.2020.1724496 Reference page. 
• Huber, B.A. & Villarreal, O. 2020. On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy 718: 1–317. DOI: 10.5852/ejt.2020.718.1101. . Reference page. 

• Villarreal M, O., García, A.F. & Kury A. B. 2021. Fine-tuning the diversity in four families of Gonyleptoidea (Arachnida, Opiliones) in Venezuela. Revista Mexicana de Biodiversidad 92(e923633): 1–22. DOI: 10.22201/ib.20078706e.2021.92.3633 Reference page. 
• Villarreal Manzanilla, O., Kury, A.B. & Colmenares, P.A. 2021. Contributions to the taxonomy of some Amazonian Stygnidae (Opiliones: Laniatores: Gonyleptoidea). Zootaxa 4984(1): 218–227. DOI: 10.11646/zootaxa.4984.1.17. . Reference page. 

• García, A.F., Kury A. B. & Villarreal M, O. 2022. Chasing the elusive core identity of Avima Roewer, 1949 (Arachnida: Opiliones: Agoristenidae). Arthropoda Selecta 31(2): 217–227. DOI: 10.15298/arthsel.31.2.10 Reference page. 

• García, A.F. & Villarreal M, O. 2023. Looks are deceiving: a cladistic analysis, three new species, and a new diagnosis of Paravima Caporiacco, 1951 (Opiliones: Agoristenidae). Arthropod Systematics & Phylogeny 81: 409–437. DOI: 10.3897/asp.81.e85571 [https://arthropod-systematics.arphahub.com/article/85571/list/1/ PDF Reference page. 
• Villarreal M, O., Ahumada-C., D. & Delgado-Santa, L. 2023. Mapping the distribution of armored harvestmen (Opiliones, Laniatores) in Colombia: updated list of species, taxonomic contributions, and insight of diversity in protected areas. Zookeys 1175ː 223–284. DOI: 10.3897/zookeys.1175.102485 Reference page. 
• Villarreal M, O., Medrano, M.A. & Kury, A.B. 2023. Unleashing Abria Sørensen, 1932: New generic rank proposal for a false Venezuelan Cynorta Koch, 1839 (Opiliones: Cosmetidae). Zoologischer Anzeiger 305: 42–51. DOI: 10.1016/j.jcz.2023.06.001 Reference page. 

• Villarreal Manzanilla, O., Ahumada-Cabarcas, D. & Flórez-Daza, Á.E. 2024. A glimpse into the diversity of Stygnus Perty, 1833: A new Colombian species and additional records from South America (Opiliones, Stygnidae). European Journal of Taxonomy 970: 203–229. DOI: 10.5852/ejt.2024.970.2759. . Reference page. 
• Villarreal Manzanilla, O., González-Gómez, J.C., Kury, A.B. & García, L.F. 2024. Peering beyond the monotypic veil: taxonomy and notes on the parental care of Neocranaus (Opiliones: Gonyleptoidea: Cranaidae). European Journal of Taxonomy 925: 292–320. DOI: 10.5852/ejt.2024.925.2471. . Reference page. 

• Colmenares, P.A.; Moreno-González, J.A.; Villarreal M, O. & Prendini, L. 2025. New genus and species of short-tailed whipscorpion (Schizomida: Hubbardiidae) from the Venezuelan Amazon. Zootaxa 5563(1): 84–95. DOI: 10.11646/zootaxa.5563.1.9 Reference page.